INTRODUCTION
Rainfed mixed crop/livestock farming is the principal production system in the highlands of Ethiopia. It is a complex system that combines crop and livestock production within the same management unit. However, the productivity of livestock in the traditional mixed farming system is very low (Haile et al., Reference Haile, Joshi, Ayalew, Tegegne and Singh2009). Livestock are also often blamed for their negative impact on the environment (Steinfeld et al., Reference Steinfeld, Gerber, Wassenaar, Castel, Rosales and de Haan2006). Improving livestock productivity while mitigating their environmental impact challenges to enhance the role of livestock in reducing poverty and improve the livelihoods of smallholder farmers in Ethiopia and across sub-Saharan Africa.
Water is a key limiting factor to livestock production in the upper Blue Nile basin of Ethiopia particularly because of the highly seasonal availability of feed resources. Livestock production is considered to be a relatively water intensive enterprise (Molden et al., Reference Molden, Oweis, Steduto, Bindraban, Hanjra and Kijne2010) regardless of the production system. Globally, there is growing concern on water allocation to the agricultural sector in the face of the on-going climate change (Zhang et al., Reference Zhang, Zwiers, Hegerl, Lambert, Gillett, Solomon, Stott and Nozawa2007) and possible competing uses (Molden et al., Reference Molden, Oweis, Steduto, Bindraban, Hanjra and Kijne2010). It is thus important to evaluate livestock productivity from a water use perspective to help comprehend strategies that can improve livestock water productivity (LWP).
LWP is a concept (Peden et al., Reference Peden, Taddesse and Haileslassie2009) that has recently received attention (Descheemaeker et al., Reference Descheemaeker, Amede and Haileselassie2010; Haileselassie et al., Reference Haileselassie, Peden, Gebreselassie, Amede and Descheemaeker2009; Molden et al., Reference Molden, Oweis, Steduto, Bindraban, Hanjra and Kijne2010) because of its usefulness as a tool to identify possible intervention options that can lead to increase livestock production with more effective use of water and with reduced impact on the environment. LWP is generally determined as a ratio of total benefits obtained from livestock per unit of water depleted in production (Peden et al., Reference Peden, Taddesse and Haileslassie2009). Previous work on LWP including the study of Haileselassie et al. (Reference Haileselassie, Peden, Gebreselassie, Amede and Descheemaeker2009) focused on assessing livestock outputs and water inputs accounted over a single study year. This approach has limitations in accounting for the portion of water used by the livestock to reach production age. The present study was, therefore, intended to determine LWP using a life cycle assessment (LCA) method to fully account for the water used by livestock from birth to farm gate. The objectives of the present study were (i) to assess LWP across different agro-ecological zones of Gumara watershed, and (ii) to highlight possible intervention options for improving LWP relevant to the study area.
MATERIALS AND METHODS
Study site
Gumara watershed was selected in the upper Blue Nile basin (Supplementary Fig. S1 available online at https://doi.org/10.1017/S0014479717000321) for undertaking the present study. The site is located in the north western part of Ethiopia having coordinates of 11°81′–11°85′ N, and 37°70′–38°02′ E. Elevation of the watershed varies from 1780 m above sea level (asl) around the entry point of Gumara River into Lake Tana to 3704 m asl towards the source of the river at the base of Guna mountain. The surface area of the watershed is 1277 km2 and it produces a mean annual flow of 1317 MCM (Abereham, Reference Abereham2013). The rainfall distribution follows a uni-modal pattern with most precipitation occurring in June–September with average annual precipitation of 1492 mm in the upper reaches and 1378 mm in the lower reaches of the watershed (Fenta, Reference Fenta2009). The landscapes of the watershed encompass various topographic features ranging from rugged rolling hills to the vast flat lands known as the Fogera plains, towards Lake Tana that is usually flooded during the wet season. The soil type of the watershed is generally classified into five categories (Fenta, Reference Fenta2009) out of which the luvisols dominate the upper reaches while the vertisols dominate the lower reaches of the watershed.
Farming systems
The prevailing agricultural production system in Gumara watershed is subsistence-based mixed crop-livestock farming. Human and livestock pressures are high relative to the resource base of the watershed and consequently out-migration takes place to other highland or lowland areas. There is an acute shortage of arable land that has led to the expansion of production on marginal and fragile lands including steep slopes (Tamene and Vlek, Reference Tamene, Vlek, Braimoh and Vlek2008).
The crops grown in the watershed area are relatively diverse depending on the agro-ecological conditions. Given the mosaic of mixed crop-livestock farming land uses occurring in the watershed, three distinct scenarios of mixed farming practice were identified. Haileselassie et al. (Reference Haileselassie, Peden, Gebreselassie, Amede and Descheemaeker2009) specified a similar division of the mixed farming system to describe the watershed and the details can be found in their report.
1. Rice/noug based farming system (RNS): It occupies the vast plain area of Fogera adjacent to Lake Tana (Fig. S1), which is warmer for most of the year. It covers approximately 20% of the watershed and is situated at altitudes between 1780 and 1850 m asl. The farming community used to rely more on livestock than on crop production for its livelihood. Livestock have become secondary to crop production after the introduction of paddy rice farming to make use of the large flooded plain area in the wet season. The community predominantly raises cattle of their own local breed known as Fogera. The high livestock density in the area is a major problem. Feed shortage becomes critical during the rainy season due to flooding and depletion of the crop residue reserves. Livestock health problems such as trypanosomosis and fasciolosis are major concerns. The present low level of livestock performance is a manifestation of the complexity of problems in livestock production in the study area.
2. Tef/finger millet based farming system (TMS): This category represents the largest part (about 60%) of the watershed. The altitude ranges between 1851 and 2400 m asl. Cool to moderately warm weather conditions prevail. The area is typically characterized by rugged terrain and rolling hills. Within a small area, wide ecological variability is found and hence a diverse range of crops is grown. Fragmented stone cover is a typical feature of the croplands. Similarly, different livestock species viz. cattle, sheep, goats, horses, donkeys and mules are kept by farm households. Equine livestock are used as pack animals for transporting agricultural goods across the prevailing rugged terrain. This farming system suffers from problems of severe soil erosion, land degradation, overutilization of communal grazing land, feed shortage, and poor health care all of which combine to produce low livestock performance.
3. Barley/potato based farming system (BPS): This system represents about 20% of the watershed area at an elevation between 2401 and 3700 m asl. The system is characterized by cool uplands with a mean daily minimum air temperature of 2.8 °C (Fenta, Reference Fenta2009). The terrain is characterized by undulating chains of mountains. Unlike the other two farming systems, the use of horses and mules for ploughing cropland is a common practice in this system. In terms of numbers, sheep are the dominant livestock species kept by farm households. Fragmented stone cover on croplands is common and similar to the TMS. Poor soil fertility as a consequence of soil erosion, land degradation and declining land holding is the major problem that poses a challenge to the farming community to sustain its livelihood. Like the other two systems, livestock shows low performance mainly due to feed shortage and health problems such as fasciolosis (particularly in sheep) on wet lowlands.
Determination of LWP
LWP was assessed at a household level across the different farming systems of Gumara watershed by taking 2–3 representative peasant associations (PAs) from each system. Again from each PA, 10 households were selected on a random basis. The sample farmers were then stratified into three wealth categories (resource-poor, -medium and -‘rich’ farmer) based on the perception of their peer colleagues in relation to the households’ relative income, herd size, land holding and annual crop harvest (Table 1). A total of 62 farmers were monitored between November 2006 and February 2008 to collect pertinent data on crop and livestock production using recruited enumerators. In the present study, LWP was determined as the ratio of the total sum of benefits (expressed in monetary value) obtained from livestock production to the amount of water depleted or degraded for producing these benefits (Peden et al., Reference Peden, Taddesse and Haileslassie2009). For determining the water used in livestock production of the study area, an LCA-based water footprint method was used to account for processes occurring from birth to exit from the farm for the whole herd as described by Koehler (Reference Koehler2008) and Peters et al. (Reference Peters, Wiedemann, Rowley and Tucker2010). The LCA method is a biophysical accounting framework used to characterize the material/energy flows underpinning specific activities and quantify their contributions to resource depletion and emissions-related environmental concerns (Pelletier et al., Reference Pelletier, Lammers, Stender and Pirog2010). In this study, it was used with a frame boundary of cradle to farm gate that enables to invoke the whole continuum of the herd life in accounting for the water use. Hence, the determination of LWP covered the period between birth and end of productive age for breeding stock, while this period was set between birth and time of offtake (slaughter or sale) for non-breeding stock.
Key features describing wealth categories of farm households in Gumara watershed area.

TLU: tropical livestock unit equivalent to 250 kg live weight; USD: US dollar.
Determination of water use
The major water requirement for livestock production is often related to production of animal feed (Peden et al., Reference Peden, Taddesse and Haileslassie2009). In all the three scenarios of the mixed farming systems, crop residues are used as a key resource for feeding livestock in the long dry season. To quantify the amount of crop residues produced by each farm household, we estimated grain and crop residue yields of cereals (barely, tef, rice, wheat, triticale, finger millet, sorghum and maize) and pulses (faba bean, field pea, chickpea, lentil and grass pea) grown by farmers in each farming system using 1×1 m quadrats. In sampling, 3 quadrats were taken from each field and all the fields were sampled for each crop grown by a household. For estimating the size of cropland allotted to each crop type, we used GPS to define field boundaries.
We partitioned the total metabolizable energy (ME) derived from the feeds consumed by each animal to account for the water portion used in meeting its maintenance requirement as indicated by NRC (2000, 2001, 2007) and Nsahlai et al. (Reference Nsahlai, Osuji and Umunna1997). The energy required for walking to allow grazing was also estimated and included in the daily maintenance component.
Biomass productivity of both private and communal pasturelands used by each household was determined by taking representative samples (5–9 samples depending on the size of the pastureland) using 0.5×0.5 m quadrats. Determining biomass of a pasture produced from a hectare of land was required in order to estimate the amount of water used to grow a given amount of pasture. The collected samples of pasture and crop residues were analysed for nutritive value. The amount of supplementary feeds, mainly crop residues, offered to animals at homesteads, was measured throughout the study period. Knowing the energy content and quantity of crop residues offered to each animal by a household, we estimated the amount of pasture intake of each animal from grazing to sustain its live weight and provide output for which the animal is kept. We estimated the amount of feed consumed by each animal over its life cycle through extrapolating the feed intake data collected, in this study, across age classes of the different livestock species. To determine the volume of water used in producing each crop type and pasture, we used the CROPWAT model that employs the Penman–Monteith equation for estimating the reference evapo-transpiration (ETo) as described by Allen et al. (Reference Allen, Pereira, Raes and Smith1998). The crop evapo-transpiration (ETc) was determined as the product of ETo by the crop coefficients using specific data (meteorological data, cropping pattern and soil data) inputs required by the CROPWAT8 computer program. Effective precipitation was estimated using the fixed percentage (potential evapo-transpiration/precipitation ratio) method in the CROPWAT model.
The water requirement (m3 ha−1) of each crop and pasture was then calculated from the accumulated ETc (mm d−1) over the complete growing period. We partitioned the total ETc of cereal and pulse crops into the grain and crop-residue components based on their respective local market values (Singh, Reference Singh, Sharma, Singh and Shah2004). We estimated drinking water consumed by different livestock species and age classes as described by FAO (1986) over the productive lifetime for the breeding stock and until time of offtake for the non-breeding animals. Water required for cleaning barns, animals, utensils and others was not estimated in this study. We did not estimate degraded water (water that cannot be recycled for use such as that mixed with acaricide in dipping animals) because of the complexity to quantify it.
Benefits from livestock
In mixed crop/livestock production systems, livestock are kept for multiple purposes. To assess the multiple benefits obtained from livestock in accounting for LWP, we quantified the various products and services rendered by the animals on the farm including the benefit from sale of animals. The estimated benefits were finally converted to monetary values based on their respective market prices to arrive at a combined monetary value. Insurance value from keeping livestock is related to the capital invested in the herd as a guarantee for meeting unexpected expenditures such as medical bills. The benefit from the insurance and financial functions is estimated to increase the value of keeping livestock by up to 7% (Bebe, Reference Bebe2003).
Livestock outputs
Livestock outputs in terms of milk production and meat production (animal offtake) were assessed until exit from a farm. Lifetime milk production was estimated through monitoring lactation yields of milking cows and carrying out a survey using questionnaires. Data on reproductive performance of sheep and goats were also collected using questionnaires. Live weight measurements of animals were taken once a month from each household across the three farming systems. Changes in herd or flock size due to birth, death or sale were monitored in each household during the study period. Breeding animals are culled at the end of their productive life by slaughter. Those offspring not required for stock replacement or for another reason are usually sold at the ages of 2 or 4 years in the case of cattle. All households indicated that they sell bulls or heifers according to two offtake scenarios (early offtake at 2 years of age and late offtake at 4 years of age), depending on circumstances at the time. Therefore, the two offtake scenarios were considered in each household to estimate their corresponding LWP. The reason why only offtake scenarios were considered as an option to improve LWP was because of the scope of the data obtained.
Draught use
Work performance (tillage, threshing and pack transport) of draught animals was monitored throughout the study period by keeping records of daily working hours and by calculating the number of working days in a year. The draught service over the productive age of draught animals was converted into monetary values by multiplying the number of work days by the local daily rate for hiring a pair of oxen or equids.
Manure and urine
Manure nutrient content under smallholder farm management conditions was averaged to be 16.1, 3.6 and 16.8 g kg−1 DM for N, P and K, respectively (Lupwayi et al., Reference Lupwayi, Girma and Haque2000). Urine contains 0.9% N and 0.5% K on a wet basis while P content is reported as trace level (FAO, 1992). We estimated the output of urine over the herd life or until offtake using a daily average rate of 31 mL kg−1 body weight (FAO, 1992) regardless of its disposal site. Manure output was estimated at a rate of 3.3 kg for cattle and 2.4 kg for small ruminant and equine per day per TLU (Haileselassie et al., Reference Haileselassie, Priess, Veldkamp and Lesschen2006). We considered the nutrient price of inorganic fertilizer (Mekonnen et al., Reference Mekonnen, Descheemaeker, Tolera and Amede2011) for estimating monetary value of manure and urine to serve as soil amelioration input. The monetary value of the dung portion used for fuel, estimated at 38% of the total output (Hawando, Reference Hawando2000), was assessed based on its market price.
Statistical analysis
The General Linear Model of SAS (2002) was employed to analyse the data. For testing effects of the independent factors on response variables, the statistical model used in the analysis is
where Yijk = response variable such as LWP, water use; µ = the overall mean, Wi = ith wealth status of smallholder farmers, Fj = jth farming system, (W*F)ij = interaction between wealth status and farming system and Eijk = error term.
The interaction effect in the model was not found to be significant (p > 0.05) in all the cases and hence was left out. A group t-test procedure was run to compare the means of LWP between early offtake and late offtake using Satterthwaite's approximation for unequal variances (SAS, 2002). A separate analysis was also carried out to test the effect of livestock species on dry matter, metabolizable energy intakes and LWP.
RESULTS
Livestock performance
Small ruminants had a lower (p < 0.01) daily dry matter intake per kg of metabolic body weight than both cattle and equine (Table 2). Hence, the ME used per kg of metabolic body weight was significantly lower (p < 0.001) in small ruminants than in cattle reflecting their lower metabolic rate. The proportion of cattle (on a weight basis) out of the total TLU kept by a household tended to inversely relate with altitude, this being manifested by a higher number of cattle in the RNS than in the BPS (Table 3). Of all the livestock outputs, larger returns (USD TLU−1 year−1) were obtained from manure used as fertilizer and sale of live animals. The return from draught power use was also considerable, amounting to about 14% of the monetary value derived from livestock. The benefit from milk production in monetary terms seemed to be among the lowest although its contribution in improving the nutritional intake of the poor rural family mainly of the children is invaluable. Monetizing the value of insurance function of the traditional livestock production by smallholder farmers appeared to raise the total livestock benefit by 6% (Table 3). Nonetheless, mortality incidence was found to reduce the total return from livestock production by 28% annually on average (Table 3).
Live weight and daily feed consumption of livestock species kept by smallholder farmers in Gumara watershed.

DM: Dry matter; ME: metabolizable energy; se: standard error; lwt: live weight; **: significant at 1%; means with different superscript letters in a column are significantly different at the indicated significance level.
Livestock holding and values of livestock outputs at a household level in three mixed farming systems.

† Number in parenthesis represents percent share of cattle in TLU on live weight basis; TLU: tropical livestock unit equivalent to 250 kg live weight; USD: US dollar; SC: represents the livestock capital kept at farm; Sale: return from sale of live animal; DP: draught power; Manr: manure; Insur: insurance value of keeping stocks; Mort loss: mortality loss; **: significant at 1%; *: significant at 5%; ns: not significant; means with different superscript letters in a column are significantly different at the indicated significance level.
Water productivity
In Gumara watershed, smallholder farmers cultivated diverse crops owing to heterogeneity in farm conditions and spreading the risk of crop failure. Combining the benefits of these crops together in monetary terms, RNS showed a higher CWP (p < 0.01) with a value of 0.46 USD m−3 of water than both BPS and TMS (Table 4). In contrast, LWP was found to be significantly lower (p < 0.01) in RNS (0.057 USD m−3 water) than in TMS (0.066USD m−3 water) or in BPS (0.066 USD m−3 water). This indicates that water requirement per kg live weight of animal increased towards lower altitude areas implying more water loss in RNS. As a result, 20% additional water input was required to sustain each kg of live weight of livestock in RNS as compared to BPS in the upper reach of the watershed. CWP was superior to LWP across the mixed farming systems of Gumara watershed. Wealth status of a household appeared to affect both crop and livestock water productivities (Table 5). Only ‘rich’ farmers were able to attain both higher CWP and LWP.
Least-squares means of crop water productivity, LWP and water used to sustain a kilogram of live weight of livestock under three different mixed farming systems.

CWP: economic crop water productivity of various crops grown by a household combined together in monetary value per unit of water input; LWP: livestock water productivity; **: significant at 1%; *: significant at 5%; means with different superscript letters in a column are significantly different at the indicated significance level.
Least squares means of CWP, LWP and water use per kg live weight of livestock by wealth status of smallholder farmers in Gumara watershed.

CWP: economic crop water productivity of various crops grown by a household combined together in monetary value per unit of water input; LWP: livestock water productivity; **: significant at 1%; *: significant at 5%; means with different superscript letters in a column are significantly different at the indicated significance level.
A cattle herd management strategy that pursued early offtake increased LWP by 28% over that of late offtake (Table 6). The amount of water used per kg live weight of the animal over its entire lifetime was much lower (reduced by more than 50%) in the early offtake scenario than in the late offtake. Contrasting the income from sale of animals per TLU, early offtake gave a higher return than late offtake.
Means of LWP, income from sale of live animal and water use to sustain a kilogram of live weight of cattle under two offtake scenarios.

LWP: livestock water productivity; **: significant at 1%.
DISCUSSION
LWP and its methodology
Water productivity is a useful concept to gauge the extent of water depleted for agricultural production and identify interventions to improve water use efficiency. This is important because water has become a very scarce resource (Molden et al., Reference Molden, Oweis, Steduto, Bindraban, Hanjra and Kijne2010) and food production is increasingly constrained by water shortage (Wallace, Reference Wallace2000). Regarding its application to livestock production, the concept of LWP as first developed by Peden et al. (Reference Peden, Taddesse and Haileslassie2009) helps to clearly understand the interaction between livestock and water including its implications for the production environment (e.g. water contamination from livestock wastes and degraded water). Estimating LWP is a complex exercise as it requires data on the performance of livestock and conversion of all livestock outputs/utilities to monetary values using contemporary price indices. In Ethiopia, smallholder farmers do not keep performance records for their animals. The challenges encountered in obtaining complete performance data of animals, on a long term basis, might have caused some degree of uncertainty to precisely determine LWP in our study. Furthermore, capturing all the intangible values of keeping livestock in the study area was challenging although we attempted to monetize insurance value of livestock (Bebe, Reference Bebe2003). In a similar study, Moll (Reference Moll2005) also highlighted the difficulty in monetizing the intangible values (accumulation of wealth, security against contingencies, display of status and others) of livestock production in sub-Saharan Africa. Hence, caution must be taken when comparing LWP values across countries or regions because of variability in production targets and in market prices of livestock outputs or services.
The methodology for quantifying the amount of water used to produce animal feeds should be refined and standardized based on the goals of livestock production. To have a realistic estimate of water depletion in livestock production, it is necessary to assess the feed intakes of different livestock species from birth to farm gate and the source of feeds in all steps of the production. Since animal feed accounts for more than 95% of the water portion required in livestock production (Peden et al., Reference Peden, Taddesse and Haileslassie2009), proper estimation of the feed consumed by livestock is a determining factor for reaching reliable LWP values. In the present study, the water input from feed consumption (through grazing and supplementary feeding) of each animal was assessed from birth to farm gate using an LCA method. Our estimation of the total consumptive water used in livestock production is subject to some uncertainty due to the absence of intake data mainly from grazing. When feed intake does not meet the energy requirement of an animal, metabolic energy is derived from animals' body fat, protein and glycogen for example to allow on-going production of milk. However, we found it difficult to have a complete account of the energy portion derived from metabolism of the stored energy that causes body weight loss of animals.
In Gumara watershed, LWP ranged from as low as 0.03 to as high as 0.1 USD m−3 water across the different agro-ecologies. These figures appear to be low compared to those of CWP (0.2–0.6 USD m−3) obtained from the same domain. The implication is that the intrinsic feature of livestock production makes the enterprise to be more water intensive. The reasons for this are complex and context specific. However, the necessity for an animal to first pass through a long growing period in its life cycle (retarded growth at early age due to poor management in the study area) to reach its productive age is worth mentioning as one of the main reasons for low LWP. Contrary to CWP, LWP was lower in RNS than in BPS or TMS (Table 4). The reason could be associated with ETc and livestock management. RNS area has a warmer climate, and hence the ETc required for both pasture and crop production was higher than in the upper catchment of the study area. The higher CWP in RNS may also have been due to its prevailing double cropping practice and better access to market linkages in relative terms.
In agreement with our results, Haileselassie et al. (Reference Haileselassie, Blummel, Clements, Descheemaeker, Amede, Samireddypalle, Sreedhar Acharya, Venkatar Radha, Ishaq, Samad, Murty and Khan2011) reported LWP estimates of 0.03–0.12 USD m−3 water for Indo-Ganga basin, and Breugel et al. (Reference Breugel, Herrero, de Steeg and Peden2010) calculated 0.01–0.13 USD m−3 water for different farming systems in the Nile Basin. Similarly, Mekonnen et al. (Reference Mekonnen, Descheemaeker, Tolera and Amede2011) and Descheemaeker et al. (Reference Descheemaeker, Amede, Haileselassie and Bossio2011) also showed comparable values of LWP for water stressed environments of northern Ethiopia with a narrower range between 0.07 and 0.09 USD m−3 water. However, the reports by Gebreselassie et al. (Reference Gebreselassie, Peden, Haileselassie and Mpairwe2009) and Haileselassie et al. (Reference Haileselassie, Peden, Gebreselassie, Amede and Descheemaeker2009) found much higher estimates of LWP ranging between 0.25 and 0.39 USD m−3 water. This divergence might have arisen from methodological differences to determine LWP. In the previous studies, the determination of LWP was limited to data of one year, which accounts for only partial input/output of the livestock production without accounting for the water inputs used in raising the animals to reach their productive age. In contrast, the present study applied the LCA method (Peters et al., Reference Peters, Wiedemann, Rowley and Tucker2010) to drive water footprints for estimating the total water depleted in livestock production by setting the limits from birth to farm gate. We also included the consumptive water accounted to those animals that exited the farm due to death.
LWP in Gumara watershed could be improved provided that appropriate interventions can be employed. In supporting this view, Molden et al. (Reference Molden, Oweis, Steduto, Bindraban, Hanjra and Kijne2010) indicated that areas where poverty is high and water productivity is low are among the priority areas where substantive increase in water productivity is possible. Furthermore, a theoretical ex-ante analysis done by Descheemaeker et al. (Reference Descheemaeker, Amede, Haileselassie and Bossio2011) for smallholder mixed crop-livestock systems of Ethiopia indicated that LWP could be tripled (to about 0.17 USD m−3 water) on the same landholding by combining different potential intervention options related to improving feed, water and livestock managements, and applying them together.
Livestock outputs and services
Manure application for sustaining soil fertility is featured as a key ecosystem component in nutrient cycling models of agricultural systems (Murwira et al., Reference Murwira, Swift, Frost, Powell, Fernandez-Rivera and Williams1995). Currently, the Ethiopian government has paid more attention than ever before on extending the use of compost (from manure and plant biomass) by smallholder farmers. In the present study, manure and draught power made up a considerable share of the total monetary value obtained from livestock component (Table 3). The higher value of draught power in TMS might be associated with the need for intensive tillage to make a fine seedbed for small seeded cereals like tef. It can, thus, be noted that the overriding reason to keep livestock as an integral part of the mixed farming system is to provide the necessary inputs required for crop production. This leads to a tradeoff between improving livestock outputs per animal and keeping more animals to support crop production (for ploughing, compacting and threshing) against a background of critical feed shortage.
In the tropics, livestock productivity is greatly reduced by poor husbandry practices coupled with animal health problems. In this study, we estimated mortality incidence at a rate of 10% in equids, 14% in cattle and 20% in small ruminants, affecting mainly young stock. Consequently, its cumulative effect on reducing the benefits (in monetary value) from livestock reached as high as 28% in the present study. Agreeing with these results, Gizaw et al. (Reference Gizaw, Tegegne, Gebremedhin and Hoekstra2010) reported mortality of sheep to be 17–26% under traditional management practices of smallholder farmers in Ethiopia. So, interventions that target improved livestock health would considerably contribute to increasing productivity of livestock and water. Feed utilization efficiency of livestock seems to correlate with LWP. Cattle had higher LWP than equids (Table 7) since they provide more products and have better digestion efficiency (Udén and Van Soest Reference Udén and Van Soest1982) when utilizing low-quality feedstuffs of the type commonly available in the study area.
Least squares means of LWP and water use to sustain a kilogram of live weight of different livestock species over their productive life time.

LWP: livestock water productivity; **: significant at 1%; means with different superscript letters in a column are significantly different at the indicated significance level.
Livestock offtake
Time of offtake of livestock appeared to have a significant influence on LWP. Prolonging the time to offtake of those animals not required for replacement to about 4 years of age or above would unnecessarily add to the cost of their maintenance resulting in lower LWP. Farmers opt to keep extra animals for longer years to exploit their uses in asset accumulation, prestigious social values, threshing crop harvests and others. However, the cost of inputs (in terms of feed consumption and water use) for keeping these extra animals beyond 2 years seemed to be relatively high although their market prices would appear to slightly rise as compared to those of their younger counterparts. Nowadays in Ethiopia, consumers are more concerned about the quality of meat and hence, livestock traders prefer buying younger animals from producers to sell them at a better price relative to their keeping cost. Promoting the offtake of younger cattle is also beneficial for encouraging the beef industry in Ethiopia to target the export market of the Middle East countries. Negassa and Jabbar (Reference Negassa and Jabbar2008) illustrated the need for incentivizing smallholder farmers to induce the supply of young animals to market and this in turn would contribute to alleviation of the overstocking problem in the highlands of Ethiopia.
CONCLUSION
The LWP determined for Gumara watershed was 0.063 USD m−3 water on average and showed wide variation between households across the farming systems. The prospect for improving LWP is considerable in light of the present growing aspiration pursued by the Ethiopian government to fuel agricultural development in the country, for instance, through keeping only productive animals, increasing pasture productivity, improving the utilization of other feed sources and linking the production goals to market demands. The interventions targeting early offtake proved to substantially increase LWP and this would contribute to ease the imbalance between the existing high livestock population and the deteriorating carrying capacity of natural pasture.
Acknowledgements
The authors would like to thank the CGIAR Challenge Program on Water and Food (CPWF) for financially supporting this study. We are grateful to the two anonymous reviewers for their immense contribution to improve the paper. We would like to thank Dr. Alan Duncun for helping us in editing the paper. Finally, our gratitude goes to the farmers in the study area for their participation and resource commitment.
SUPPLEMENTARY MATERIAL
For supplementary material for this article, please visit https://doi.org/10.1017/S0014479717000321.






