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THE CURRENT STATUS OF THE SPECIES HITHERTO ASSIGNED TO HENCKELIA (GESNERIACEAE)

Published online by Cambridge University Press:  18 October 2013

D. J. Middleton ,
A. Weber ,
T. L. Yao ,
S. Sontag  and
M. Möller
D. J. Middleton
Affiliation:
Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh EH3 5LR, Scotland, UK. E-mail for correspondence: d.middleton@rbge.ac.uk
A. Weber
Affiliation:
University of Vienna, Faculty Centre of Biodiversity, Department of Structural and Functional Botany, Rennweg 14, 1030 Vienna, Austria.
T. L. Yao
Affiliation:
Forest Research Institute Malaysia, 52109 Kepong, Selangor, Malaysia.
S. Sontag
Affiliation:
University of Vienna, Faculty Centre of Biodiversity, Department of Structural and Functional Botany, Rennweg 14, 1030 Vienna, Austria.
M. Möller
Affiliation:
Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh EH3 5LR, Scotland, UK. E-mail for correspondence: d.middleton@rbge.ac.uk

Abstract

Following recent molecular phylogenetic studies in Old World Gesneriaceae the nomenclatural implications for names in Henckelia are examined. New combinations are made in Codonoboea and Loxocarpus to account for species now excluded from Henckelia. A list is presented in which the current position of all species hitherto assigned to Henckelia is given, including the new combination Henckelia rotundata (Barnett) D.J.Middleton & Mich.Möller. A new combination in Oreocharis is made.

Keywords

CodonoboeaGesneriaceaeHenckeliaLoxocarpusnew combinations

Information

Type
Articles
Information
Edinburgh Journal of Botany , Volume 70 , Issue 3 , November 2013 , pp. 385 - 404
Copyright
Copyright © Trustees of the Royal Botanic Garden Edinburgh 2013 

Introduction

para1In recent years there has been a large number of changes in generic delimitation in Gesneriaceae as a result of both detailed morphological study and insights from molecular phylogenetic analyses (Möller et al., Reference Möller, Pfosser, Jang, Mayer, Clark and Hollingsworth2009, Reference Möller, Forrest, Wei and Weber2011a, Reference Möller, Middleton, Nishii, Wei, Sontag and Weber2011b; Wei et al., Reference Wei, Wen, Chen, Shui and Möller2010; Puglisi et al., Reference Puglisi, Middleton, Triboun and Möller2011; Wang et al., Reference Wang, Mao, Liu, Li, Dong, Li and Smith2011; Weber et al., Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a, Reference Weber, Wei, Puglisi, Mayer and Möller2011b, Reference Weber, Wei, Sontag and Möller2011c). The study by Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a) was ostensibly to clarify relationships in the heterogeneous genus Chirita Buch.-Ham. ex D.Don but the conclusions also had significant ramifications in other genera, particularly in Henckelia Spreng. The objective of the current paper is to clarify the nomenclatural consequences of changes in the delimitation of Henckelia.

Henckelia was resurrected from synonymy of Didymocarpus Wall. by Weber & Burtt (1998). It encompassed the species formerly placed in Didymocarpus sect. Orthoboea (South India, Sri Lanka), the species of all (but one: Didymocarpus sect. Elati Ridl.) sections of Didymocarpus from Peninsular Thailand and Malesia, as well as the Malesian genera Codonoboea Ridl., Loxocarpus R.Br., and Platyadenia B.L.Burtt, totalling about 180 species. The first molecular phylogenetic studies to include a range of Henckelia species confirmed that these species indeed did not form a monophyletic group with Didymocarpus but also began to suggest that the newly defined Henckelia might also not be monophyletic (Möller et al., Reference Möller, Pfosser, Jang, Mayer, Clark and Hollingsworth2009). Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a) showed conclusively that the South Indian and Sri Lankan species of Henckelia formed a monophyletic group with most species of Chirita sect. Chirita but that some species of Chirita sect. Chirita and the other sections in the genus were only distantly related to these. The type species of Henckelia, H. incana (Vahl) Spreng., is from South India. Henckelia has priority over Chirita and the nomenclatural consequences for all species previously assigned to Chirita were presented in Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a). The same study also showed that all the Malesian species of Henckelia, by far the majority of species in the genus, did not form a monophyletic group with the South Indian/Sri Lankan species. The focus in Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a) was on the fate of species assigned to Chirita but it was acknowledged there that the results and conclusions had huge implications for our understanding of the genus Henckelia. Henckelia is still recognised but with a delimitation considerably different from that used in a number of other recent works (Weber & Burtt, 1998; Vitek et al., Reference Vitek, Weber and Burtt2000; Weber, Reference Weber, Kubitzki and Kadereit2004; Banka & Kiew, Reference Banka and Kiew2009; Kiew, Reference Kiew2009). Before Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a) there were approximately 180 species of Henckelia and afterwards about 56 species. However, only 14 species, namely those of Henckelia sect. Henckelia (formerly Didymocarpus sect. Orthoboea Benth.) from South India and Sri Lanka, are common to both delimitations.

Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a) noted that the correct generic name for most of the southern Thai and Malesian species formerly placed in Henckelia is Codonoboea. The species of Henckelia sections Didymanthus (C.B.Clarke) A.Weber & B.L.Burtt, Heteroboea (Benth.) A.Weber & B.L.Burtt and Glossadenia A.Weber & B.L.Burtt (Weber & Burtt, 1998) consistently form a well-supported monophyletic group and can confidently all be assigned to Codonoboea. Kiew & Lim (Reference Kiew and Lim2011) made combinations for all the Peninsular Malaysian taxa and Kartonegoro (Reference Kartonegoro2012) has made a combination for the single species known from Sulawesi. In addition Rafidah et al. (Reference Rafidah, Kiew and Haron2011) have made the combination Codonoboea elata (Ridl.) Rafidah for the misplaced Chirita elata Ridl. Below we make combinations in Codonoboea for the remaining species in this group from Peninsular Thailand and the rest of Malesia. We acknowledge that considerably more basic taxonomic work is necessary in these areas to properly define the species but we consider it essential that they have names in Codonoboea pending that work so as to avoid conflicting nomenclatural frameworks.

A smaller number of species in Thailand and Malesia which were excluded from Henckelia by Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a) did not group with those species now in Codonoboea. These correspond to the species of Loxocarpus that had been synonymised into Henckelia by Weber & Burtt (1998). The species of Loxocarpus are found in the Asian twisted-fruited clade (using the terminology of Möller et al., Reference Möller, Pfosser, Jang, Mayer, Clark and Hollingsworth2009) but doubts remain as to whether the species form a monophyletic group (Yao, Reference Yao2012). However, given that Loxocarpus was explicitly excluded from the newly defined Henckelia by Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a) the species must revert to Loxocarpus and await more research on their relationships. The species have recently been revised by Yao (Reference Yao2012) and 20 named species are presently recognised. Most of these 20 species already have combinations in Loxocarpus but below we make new combinations where necessary.

We make new combinations only for those taxa which, to the best of our knowledge, are currently recognised. In the list below and in the list of all names ever published in Henckelia in Appendix 1 we largely follow taxon delimitation in Weber & Burtt (1998), Vitek et al. (Reference Vitek, Weber and Burtt2000), Weber et al. (Reference Weber, Burtt and Vitek2000), Burtt (Reference Burtt2001), Kiew & Lim (Reference Kiew and Lim2011) and Yao (Reference Yao2012).

We do not make combinations for the section names in Henckelia which were recognised by Weber & Burtt (1998) and which now belong to Codonoboea: sect. Didymanthus, sect. Heteroboea, and sect. Glossadenia. We abstain from making the formal transfers pending further research and simply list the appropriate sections hitherto described in Didymocarpus and Henckelia in the synonymy of Codonoboea.

A new combination in Oreocharis is made in Appendix 2.

Nomenclatural Consequences

Codonoboea

  • plantnameCodonoboea Ridl., Fl. Malay Penins. 2: 533 (1923). – Didymocarpus sect. Codonoboea (Ridl.) Kiew, Blumea 35: 171 (1990). – Type: Codonoboea leucocodon (Ridl.) Ridl., designated by Ivanina (Reference Ivanina1967).

  • synsDidymocarpus sect. Heteroboea Benth. in Benth. & Hook.f., Gen. Pl. 2: 1022 (1876). – Henckelia sect. Heteroboea (Benth.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 334 (1998 [‘1997’]). – Type: Didymocarpus crinitus (Jack) Spreng. (≡ AQ1: Typesetter: Please note special symbol which occurs in this paper ('is equivalent to')Codonoboea crinita (Jack) C.L.Lim).

  • synsDidymocarpus sect. Didymanthus C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 82 (1883). – Henckelia sect. Didymanthus (C.B.Clarke) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 334 (1998 [‘1997’]). – Type: Didymocarpus serratus R.Br. (≡ Codonoboea serrata (R.Br.) A.Weber).

  • synsDidymocarpus sect. Kompsoboea C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 87 (1883). – Type: Didymocarpus kompsoboea C.B.Clarke (≡ Codonoboea kompsoboea (C.B.Clarke) A.Weber).

  • synsDidymocarpus sect. Salicini Ridl., J. Linn. Soc. Bot. 32: 514 (1896). – Type: Didymocarpus salicinus Ridl. (≡ Codonoboea salicina (Ridl.) C.L.Lim).

  • synsDidymocarpus sect. Reptantes Ridl., J. Straits Branch Roy. Asiat. Soc. 44: 29 (1905). – Type: Didymocarpus reptans Jack (≡ Codonoboea reptans (Jack) C.L.Lim).

  • synsParaboea sect. Campanulatae Ridl., J. Straits Branch Roy. Asiat. Soc. 44: 63 (1905). – Type: Paraboea campanulata Ridl. (= Codonoboea hirta (Ridl.) Kiew).

  • synsDidymocarpus sect. Boeopsis Ridl., J. Straits Branch Roy. Asiat. Soc. 49: 22 (1907). – Type: Didymocarpus heterophyllus Ridl. (≡ Codonoboea heterophylla (Ridl.) C.L.Lim).

  • synsDidymocarpus sect. Pectinati Ridl., Fl. Malay Penins. 2: 508 (1923). – Type: Didymocarpus pectinatus Oliv. (≡ Codonoboea pectinata (Oliv.) Kiew).

  • synsPlatyadenia B.L.Burtt, Notes Roy. Bot. Gard. Edinburgh 31: 51 (1971). – Type: Platyadenia descendens B.L.Burtt (≡ Codonoboea descendens (B.L.Burtt) A.Weber).

  • synsDidymocarpus sect. Venusti Kiew, Malay. Nat. J. 48: 203 (1995) [‘Venustus’]. – Type: Didymocarpus venustus Ridl. (≡ Codonoboea venusta (Ridl.) Kiew).

  • synsHenckelia sect. Glossadenia A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 335 (1998 [‘1997’]). – Type: Henckelia flavobrunnea (Ridl.) A.Weber (≡ Codonoboea flavobrunnea (Ridl.) Kiew).

para1Perennial herbs, caulescent or forming a rosette, occasionally creeping; usually somewhat woody. Leaves opposite or alternate, petiolate to sessile. Inflorescences axillary, one- to few- to many-flowered, pedunculate, single-flowered or cymose, then flowers in pairs; flowers 5-merous. Calyx of 5 lobes divided to the base. Corolla consisting of a tube and 5 spreading lobes, 2-lipped with a 3-lobed lower lip, which is generally longer, and a 2-lobed upper lip. Fertile stamens 2, anthers adnate face to face. Disc a flat-topped or lobed ring, sometimes not a complete ring or lacking. Ovary cylindric, style narrower, stigma peltate or clavate. Capsules plagiotropic, usually narrow and long, dehiscing dorsally to form a gutter.

para1About 120 species from southern Thailand and throughout Malesia. The Peninsular Malaysian species are enumerated in Kiew & Lim (Reference Kiew and Lim2011) and the single Sulawesi species by Kartonegoro (Reference Kartonegoro2012). Below are new combinations for the remaining species. We have not indicated the type of type (holotype, lectotype, isotype, etc.) for all names because for those that possibly lack a holotype, or have not yet been lectotypifed, this should be done as part of a detailed revision of the taxa.

  • plantnameCodonoboea alternifolia (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus alternifolius C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 106 (1883). – Paraboea alternifolia (C.B.Clarke) B.L.Burtt, Notes Roy. Bot. Gard. Edinburgh 24: 46 (1962). – Henckelia alternifolia (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 339 (1998 [‘1997’]). – Type: W Borneo, Kalimantan, Sebalouw, near Sambas, 15–19 x 1874, Teysmann 10779 (holo FI!). – Distribution: Borneo.

  • plantnameCodonoboea amoena (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus amoenus C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 87 (1883). – Henckelia amoena (C.B.Clarke) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Type: Borneo borealis, 1877–78, Burbidge s.n. (BM, K). – Distribution: Borneo.

  • plantnameCodonoboea angustifolia (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus angustifolius C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 89 (1883). – Henckelia angustifolia (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 339 (1998 [‘1997’]). – Type: Borneo, Sarawak, in monte Mattang, Beccari 3629 (FI-B, K). – Distribution: Borneo.

  • plantnameCodonoboea appressipilosa (B.L.Burtt) D.J.Middleton, comb. nov. – Henckelia appressipilosa B.L.Burtt, Thai Forest Bull., Bot. 29: 98 (2001). – Type: Thailand, Narathiwat, Sungai Kolok, Nikhom Waeng, 2 iii 1974, K. Larsen & S. Larsen 32803 (holo AAU). – Distribution: Thailand.

  • plantnameCodonoboea bakoensis (B.L.Burtt) A.Weber, comb. nov. – Didymocarpus bakoensis B.L.Burtt, Notes Roy. Bot. Gard. Edinburgh 36: 152 (1978). – Henckelia bakoensis (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Type: Sarawak, First Division, Bako National Park, Telok Asam, 17 v 1962, Burtt & Woods 1839 (holo E; iso SAR). – Distribution: Borneo.

  • plantnameCodonoboea bakoensis (B.L.Burtt) A.Weber var. tenuior (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus leptocalyx C.B.Clarke var. tenuior C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 91 (1883). – Henckelia bakoensis (B.L.Burtt) B.L.Burtt var. tenuior (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Type: Borneo, prope Kuteing [Kuching], Beccari (holo FI-B). – Distribution: Borneo.

  • plantnameCodonoboea battamensis (Ridl.) A.Weber, comb. nov. – Didymocarpus battamensis Ridl., J. Straits Branch Roy. Asiat. Soc. 49: 22 (1908). – Henckelia battamensis (Ridl.) B.L.Burtt, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Type: Pulau Battam, an island south of Singapore, ix 1905, Kloss s.n. (SING). – Distribution: Batam Island (Indonesia).

  • plantnameCodonoboea beccarii (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus beccarii C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 89 (1883). – Henckelia beccarii (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Syntypes: Sarawak, in colle Sakarang, Beccari 3884 (FI-B); in colle Cumpang, v 1867, Beccari 3442 (FI-B, K); Kapuas, Teysmann in herb. Beccari 8441 (FI-B). – Distribution: Borneo.

  • plantnameCodonoboea bullata (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus bullatus C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 92 (1883). – Henckelia bullata (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 339 (1998 [‘1997’]). – Syntypes: Borneo, Sarawak, 750 m, Lobb s.n. (K); Beccari 2108 (FI-B). – Distribution: Borneo.

  • plantnameCodonoboea corniculata (Jack) D.J.Middleton, comb. nov. – Didymocarpus corniculatus Jack, Malayan Misc. I, 2: 4 (1820). – Henckelia corniculata (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 13 (1827). – Type: Sumatra, Tappanooly, Jack s.n. (G). – Distribution: Sumatra.

  • plantnameCodonoboea corrugata (Mendum) D.J.Middleton, comb. nov. – Henckelia corrugata Mendum, Edinburgh J. Bot. 58: 439 (2001). – Type: Philippines, Palawan, side of ravine above Camp 3 towards Cleopatra’s Needle, 940 m, 22 i 1998, Mendum et al. 25558 (holo PNH; iso E). – Distribution: Philippines (Palawan).

  • plantnameCodonoboea crenata (Baker) A.Weber, comb. nov. – Didymocarpus crenatus Baker, Kew Bull. 1896: 25 (1896). – Henckelia crenata (Baker) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Type: Borneo, Sabah, Sandakan, iv 1895, Creagh (holo K). – Distribution: Borneo.

  • plantnameCodonoboea dentata (Ridl.) A.Weber, comb. nov. – Didymocarpus dentatus Ridl., J. Linn. Soc. 32: 515 (1896). – Henckelia dentata (Ridl.) B.L.Burtt, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Type: Indonesia, Lingga, 1893, Hullett s.n. (SING). – Distribution: Lingga Islands (Indonesia).

  • plantnameCodonoboea descendens (B.L.Burtt) A.Weber, comb. nov. – Platyadenia descendens B.L.Burtt, Notes Roy. Bot. Gard. Edinburgh 31: 51 (1971). – Henckelia descendens (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Type: Sarawak, First Division, Bako National Park, Telok Delima, 19 v 1962, Burtt & Woods B.1856 (holo E; iso SAR). – Distribution: Borneo.

  • plantnameCodonoboea elegans (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus elegans C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 88 (1883). – Henckelia elegans (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Type: Borneo, 1853, Lobb s.n. (holo K). – Distribution: Borneo.

  • plantnameCodonoboea filicalyx (B.L.Burtt) D.J.Middleton, comb. nov. – Henckelia filicalyx B.L.Burtt, Thai Forest Bull., Bot. 29: 95 (2001). – Type: Thailand, Narathiwat, Nikhom Waeng, 4 iii 1974, K. Larsen & S. Larsen 32914 (holo AAU; iso E). – Distribution: Thailand.

  • plantnameCodonoboea follicularis (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus follicularis C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 98 (1883). – Henckelia follicularis (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 345 (1998 [‘1997’]). – Type: Borneo, Karimata, Teysmann in herb. Beccari 11218 (FI-B; photo E). – Distribution: Borneo.

  • plantnameCodonoboea gracilipes (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus gracilipes C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 97 (1883). – Henckelia gracilipes (C.B.Clarke) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Type: Borneo, Sarawak, Beccari 3229 (FI-B, G, K, M, P, W). – Distribution: Borneo.

  • plantnameCodonoboea koerperi (B.L.Burtt) A.Weber, comb. nov. – Didymocarpus koerperi B.L.Burtt, Notes Roy. Bot. Gard. Edinburgh 47: 236 (1990). – Henckelia koerperi (B.L.Burtt) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 347 (1998 [‘1997’]). – Type: Cultivated plant originally from Sumatra, Pajakumbuh, Harau valley, c.800 m, Körper s.n. (holo E; iso M, WU). – Distribution: Sumatra.

  • plantnameCodonoboea kolokensis (B.L.Burtt) D.J.Middleton, comb. nov. – Henckelia kolokensis B.L.Burtt, Thai Forest Bull., Bot. 29: 95 (2001). – Type: Thailand, Narathiwat, Sungai Kolok, Nikhom Waeng, 4 iii 1974, K. Larsen & S. Larsen 32913 (holo AAU; iso E). – Distribution: Thailand.

  • plantnameCodonoboea kompsoboea (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus kompsoboea C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 92, tab. 10 (1883). – Henckelia kompsoboea (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 347 (1998 [‘1997’]). – Type: Borneo, ‘Sarawak, in colle Kumpang’, Beccari 3440 (holo FI-B). – Distribution: Borneo.

  • plantnameCodonoboea lanceolata (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus lanceolatus C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 89 (1883). – Henckelia lanceolata (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Type: ‘Borneo boreali-orientalis; in montibus’, Lobb s.n. (holo K). – Distribution: Borneo.

  • plantnameCodonoboea leptocalyx (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus leptocalyx C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 91 (1883). – Henckelia leptocalyx (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Syntypes: Borneo, Sarawak, 750 m, Beccari 1794 (FI-B, G-DC, K, P); Lobb s.n. (K). – Distribution: Borneo.

  • plantnameCodonoboea murutorum (B.L.Burtt) A.Weber, comb. nov. – Didymocarpus murutorum B.L.Burtt, Notes Roy. Bot. Gard. Edinburgh 36: 153 (1978). – Henckelia murutorum (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Type: Sarawak, Fifth Division, route from Bakelalan to Gunung Murud above Sungei Konap, c.1320 m, 23 ix 1967, Burtt & Martin B.5172 (holo E; iso SAR). – Distribution: Borneo.

  • plantnameCodonoboea myricifolia (Ridl.) A.Weber, comb. nov. – Didymocarpus myricifolius Ridl., J. Straits Branch Roy. Asiat. Soc. 44: 53 (1905). – Henckelia myricifolia (Ridl.) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Type: Borneo, Sabah, Bongaya, Labuk Bay, Ridley s.n. (SING). – Distribution: Borneo.

  • plantnameCodonoboea nervosa (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus nervosus C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 95 (1883). – Henckelia nervosa (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Type: Borneo, Sarawak, in colle Sakarang, Beccari 3889 (holo FI-B). – Distribution: Borneo.

  • plantnameCodonoboea pagonensis (B.L.Burtt) A.Weber, comb. nov. – Henckelia pagonensis B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 437 (1996). – Type: Brunei, Temburong, Bukit Pagon, east ridge, 1470 m, 19 vii 1990, K.M. Wong 1784 (holo BRUN). – Distribution: Borneo.

  • plantnameCodonoboea pleuropogon (B.L.Burtt) A.Weber, comb. nov. – Didymocarpus pleuropogon B.L.Burtt, Notes Roy. Bot. Gard. Edinburgh 31: 44 (1971). – Henckelia pleuropogon (B.L.Burtt) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Type: Cultivated plant grown on from Burtt & Martin B.5050, originally from Sarawak, Third Division, SE Hose Mts., hill E of overhang at Bukit Semako, 16 viii 1967, Royal Botanic Garden Edinburgh cultivated specimen C.5639 (holo E). – Distribution: Borneo.

  • plantnameCodonoboea porphyrea (B.L.Burtt) D.J.Middleton, comb. nov. – Henckelia porphyrea B.L.Burtt, Thai Forest Bull., Bot. 29: 96 (2001). – Type: Thailand, Ranong, Khao Pota Luang Kaeo, 22 vi 1974, Geesink et al. 7417 (holo BKF; iso AAU). – Distribution: Thailand.

  • plantnameCodonoboea punctata (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus punctatus C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 90 (1883). – Henckelia punctata (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 353 (1998 [‘1997’]). – Type: ‘Borneo boreali-orientalis, in ripis fluminum’, Lobb s.n. (holo K). – Distribution: Borneo.

  • plantnameCodonoboea racemosa (Jack) A.Weber, comb. nov. – Didymocarpus racemosus Jack, Trans. Linn. Soc. 14: 34 (1823). – Henckelia racemosa (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 13 (1827). – Type: Sumatra, west coast, at Tappanooly, Jack s.n. (probably lost). – Distribution: Sumatra.

  • plantnameCodonoboea reticulosa (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus reticulosus C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 90 (1883). – Henckelia reticulosa (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 354 (1998 [‘1997’]). – Syntypes: Borneo, Sarawak, 750 m, 1865–68, Beccari 1713 (FI-B); 1865–68, Beccari 2067 (FI-B, G-DC, K, P); Lobb (K). – Distribution: Borneo.

  • plantnameCodonoboea scabrinervia (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus scabrinervius C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 91 (1883). – Henckelia scabrinervia (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 355 (1998 [‘1997’]). – Syntypes: Borneo, ‘Sarawak, in monte Poe, alt. 1400 m’, Beccari 2423 (FI-B); Montrado, Teysmann 10883 (FI-B, L). – Distribution: Borneo.

  • plantnameCodonoboea serrata (R.Br.) A.Weber, comb. nov. – Didymocarpus serratus R.Br., Cyrtandreae 119 (1839). – Henckelia serrata (R.Br.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Type: Sumatra?, Horsfield s.n. (BM). – Distribution: probably Sumatra.

  • plantnameCodonoboea simplex (Kraenzl.) A.Weber, comb. nov. – Didymocarpus simplex Kraenzl., Mitt. Inst. Allg. Bot. Hamburg 7: 88 (1927). – Henckelia simplex (Kraenzl.) B.L.Burtt, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Type: West-Borneo [Kalimantan], ‘zwischen dem Bukit Raja und dem Bukit Menabai, um 1000 m’, 25 xii 1924, Winkler 1058 (HBG; photo E). – Distribution: Borneo.

  • plantnameCodonoboea teres (C.B.Clarke) A.Weber, comb. nov. – Didymocarpus teres C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 88 (1883). – Henckelia teres (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 357 (1998 [‘1997’]). – Syntypes: Borneo, ‘Sarawak, prope Kuteing’ [Kuching], Beccari 385 (FI-B, K); Beccari 1353 (FI-B). – Distribution: Borneo.

  • plantnameCodonoboea virginea (B.L.Burtt) A.Weber, comb. nov. – Didymocarpus virgineus B.L.Burtt, Notes Roy. Bot. Gard. Edinburgh 36: 154 (1978). – Henckelia virginea (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 358 (1998 [‘1997’]). – Type: Borneo, Sarawak, Fifth Division, route from Bakelelan to Gunung Murud, ridge above Sungai Bor, c.4°N, 115°38′E, c.1320 m, 25 ix 1967, Burtt & Martin B.5203 (holo E; iso SAR). – Distribution: Borneo.

  • plantnameCodonoboea vulcanica (Ridl.) A.Weber, comb. nov. – Didymocarpus vulcanicus Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 80 (1923). – Henckelia vulcanica (Ridl.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 359 (1998 [‘1997’]). – Type: Sumatra, Bandar Bharu, Gunong Sibayak, 6 viii 1921, Nur 7314 (K). – Distribution: Sumatra.

  • plantnameCodonoboea woodii (Merr.) A.Weber, comb. nov. – Didymocarpus woodii Merr., Philipp. J. Sci. 26: 489 (1925). – Henckelia woodii (Merr.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 359 (1998 [‘1997’]). – Type: Philippines, Palawan, Malampaya Bay, 8 x 1922, Merrill 11587 (G, K, L, P, PNH, US, W). – Distribution: Philippines.

Loxocarpus

  • plantnameLoxocarpus R.Br., Cyrtandreae 120 (1839). – Didymocarpus Wall. sect. Loxocarpus (R.Br.) Benth. in Benth. & Hook.f., Gen. Pl. 2: 1022 (1876). – Roettlera Vahl sect. Loxocarpus (R.Br.) Fritsch in Nat. Pflanzenfam. 4, 36: 147 (1894). – Henckelia Spreng. sect. Loxocarpus (R.Br.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 334 (1998). – Type: Loxocarpus incanus R.Br.

para1Rosette plants with or without rootstock, sometimes creeping, rarely caulescent. Indumentum usually of a layer of silky hairs, generally whitish, sometimes buff, usually dense, rarely sparse on lamina upper surface. Leaves petiolate. Inflorescence a reduced cyme with a single flower, or cymose, pedunculate. Flowers 5-merous. Calyx lobes free almost to base. Corolla usually campanulate or flat-faced, rarely shortly tubular; lobes more or less isomorphic but lower three lobes usually distinguishably larger than upper two lobes, upper lobes rarely fused. Disc usually absent, sometimes vestigial. Stamens 2; anthers coherent, usually included within corolla tube in campanulate and shortly tubular flowers or exposed in flat-faced flowers. Ovary conical, usually short and thick at base, sometimes slender; stigma punctiform, sometimes capitate and rarely peltate. Capsules plagiocarpic, often ovoid or conical, sometimes slender and gutter shaped, usually straight or slightly curved, rarely corniculate, dehiscing dorsally to form a splash cup or gutter.

para1More than 20 species in southern Thailand and western Malesia.

  • plantnameLoxocarpus coodei (B.L.Burtt) T.L.Yao, comb. nov. – Henckelia coodei B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 437 (1996). – Type: Brunei, Temburong, Temburong River at Wong Nguan rapids, 120 m, 5 iii 1990, Coode 6617 (holo K; iso BRUN).

  • plantnameLoxocarpus incanus R.Br. var. sekayuensis (Banka & Kiew) T.L.Yao, comb. et stat. nov. – Henckelia sekayuensis Banka & Kiew, Edinburgh J. Bot. 66: 254 (2009). – Type: Peninsular Malaysia, Terengganu, Sekayu Recreational Forest, 29 viii 1986, Anthonysamy SA 638 (holo KEP).

  • plantnameLoxocarpus sericiflavus (Kiew & Banka) T.L.Yao, comb. nov. – Henckelia sericiflava Kiew & Banka, Edinburgh J. Bot. 66: 18 (2009). – Type: Peninsular Malaysia, Johor, Gunung Panti, 5 xii 1936, Corner SFN 32538 (holo SING; iso SAR).

  • plantnameLoxocarpus taeniophyllus (B.L.Burtt) T.L.Yao, comb. nov. – Henckelia taeniophylla B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Type: Brunei, Temburong, Bangar, northern slope of Bkt. Bangar, 10–100 m, 18 i 1964, Hotta 13263 (holo KYO; iso E).

  • plantnameLoxocarpus violoides (C.B.Clarke) T.L.Yao, comb. nov. – Didymocarpus violoides C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 97 (1883). – Roettlera violoides (C.B.Clarke) Kuntze, Rev. Gen. Pl. 2: 477 (1891). – Henckelia violoides (C.B.Clarke) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Type: NE of Borneo, Lobong Peak, 5000 ft, Lobb s.n. (holo K).

Appendix 1

Appendix 1 para1Present placement of species referred to Henckelia. Current name in bold. For further nomenclatural details, where necessary, see Weber & Burtt (1998), Weber et al. (Reference Weber, Burtt and Vitek2000), Vitek et al. (Reference Vitek, Weber and Burtt2000), Burtt (Reference Burtt2001), Kiew & Lim (Reference Kiew and Lim2011) and Yao (2012AQ2: Typesetter: Please set the rest of Appendix 1 in 9/11pt with hanging indent, as shown (the style is plant name 9/11pt but without space before except for the first one)).

  • plantnameHenckelia adenocalyx (Chatterjee) D.J.Middleton & Mich.Möller, Taxon 60(3): 774 (2011).

  • plantnameHenckelia alata (Roxb.) A.Dietr., Sp. Pl., ed. 6, 1: 573 (1831). – Torenia alata (Roxb.) Benth.

  • plantnameHenckelia alba (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 338 (1998 [‘1997’]). – Codonoboea alba (Ridl.) C.L.Lim

  • plantnameHenckelia albomarginata (Hemsl.) A.Weber, Beitr. Biol. Pflanzen 70: 338 (1998 [‘1997’]). – Codonoboea albomarginata (Hemsl.) Kiew

  • plantnameHenckelia alternans (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 338 (1998 [‘1997’]). – Codonoboea alternans (Ridl.) Kiew

  • plantnameHenckelia alternifolia (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 339 (1998 [‘1997’]). – Codonoboea alternifolia (C.B.Clarke) A.Weber

  • plantnameHenckelia amoena (C.B.Clarke) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Codonoboea amoena (C.B.Clarke) A.Weber

  • plantnameHenckelia anachoreta (Hance) D.J.Middleton & Mich.Möller, Taxon 60(3): 774 (2011).

  • plantnameHenckelia angusta (C.B.Clarke) D.J.Middleton & Mich.Möller, Taxon 60(3): 774 (2011).

  • plantnameHenckelia angustifolia (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 339 (1998 [‘1997’]). – Codonoboea angustifolia (C.B.Clarke) A.Weber

  • plantnameHenckelia anthonyi (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 339 (1998 [‘1997’]). – Codonoboea anthonyi (Kiew) C.L.Lim

  • plantnameHenckelia anthonysamyi Banka, Edinburgh J. Bot. 66(2): 249 (2009). – Synonym of Loxocarpus sericiflavus (Kiew & Banka) T.L.Yao

  • plantnameHenckelia appressipilosa B.L.Burtt, Thai Forest Bull., Bot. 29: 98 (2001). – Codonoboea appressipilosa (B.L.Burtt) D.J.Middleton

  • plantnameHenckelia argentea (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 339 (1998 [‘1997’]). – Loxocarpus argenteus B.L.Burtt

  • plantnameHenckelia aromatica (D.Don) Spreng., Syst. Veg., ed. 16, 4(2): 13 (1827). – Didymocarpus aromaticus D.Don

  • plantnameHenckelia ascendens (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 339 (1998 [‘1997’]). – Codonoboea ascendens (Ridl.) C.L.Lim

  • plantnameHenckelia atrosanguinea (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Codonoboea atrosanguinea (Ridl.) C.L.Lim

  • plantnameHenckelia auriculata (J.M.Li & S.X.Zhu) D.J.Middleton & Mich.Möller, Taxon 60(3): 774 (2011).

  • plantnameHenckelia bakoensis (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Codonoboea bakoensis (B.L.Burtt) A.Weber

  • plantnameHenckelia bakoensis var. tenuior (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Codonoboea bakoensis (B.L.Burtt) A.Weber var. tenuior (C.B.Clarke) A.Weber

  • plantnameHenckelia barbata (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 13 (1827). = Liebigia speciosa (Blume) Endl.

  • plantnameHenckelia battamensis (Ridl.) B.L.Burtt, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Codonoboea battamensis (Ridl.) A.Weber

  • plantnameHenckelia beccarii (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Codonoboea beccarii (C.B.Clarke) A.Weber

  • plantnameHenckelia bifolia (D.Don) A.Dietr., Sp. Pl., ed. 6, 1: 574 (1831).

  • plantnameHenckelia bombycina (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Codonoboea bombycina (Ridl.) C.L.Lim

  • plantnameHenckelia breviflora (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 340 (1998 [‘1997’]). – Codonoboea breviflora (Ridl.) Kiew

  • plantnameHenckelia briggsioides (W.T.Wang) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia browniana A.Weber, Beitr. Biol. Pflanzen 70: 341 (1998 [‘1997’]). – Loxocarpus incanus R.Br.

  • plantnameHenckelia bullata (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 341 (1998 [‘1997’]). – Codonoboea bullata (C.B.Clarke) A.Weber

  • plantnameHenckelia burttii D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia caelestis (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 341 (1998 [‘1997’]). – Codonoboea caelestis Ridl.

  • plantnameHenckelia caerulea (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 341 (1998 [‘1997’]). – Loxocarpus caeruleus (Ridl.) Ridl.

  • plantnameHenckelia calcarea (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 341 (1998 [‘1997’]). – Codonoboea calcarea (Ridl.) Kiew

  • plantnameHenckelia calva (C.B.Clarke) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia capensis A.Braun ex C.B.Clarke in A.DC. & C.DC., Monogr. Phan. 5(1): 151 (1883), nom. inval.

  • plantnameHenckelia castaneifolia (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 341 (1998 [‘1997’]). – Codonoboea castaneifolia (Ridl.) Kiew

  • plantnameHenckelia caulescens (B.L.Burtt) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 342 (1998 [‘1997’]). – Loxocarpus caulescens B.L.Burtt

  • plantnameHenckelia ceratoscyphus (B.L.Burtt) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia cinerea (D.Don) Spreng., Syst. Veg., ed. 16, 4(2): 14 (1827). – Didymocarpus cinereus D.Don

  • plantnameHenckelia codonion (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 342 (1998 [‘1997’]). – Codonoboea codonion (Kiew) C.L.Lim

  • plantnameHenckelia communis (Gardner) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia conicapsularis (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 342 (1998 [‘1997’]). – Loxocarpus conicapsularis (C.B.Clarke) B.L.Burtt

  • plantnameHenckelia coodei B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 437 (1996). – Loxocarpus coodei (B.L.Burtt) T.L.Yao

  • plantnameHenckelia corneri (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 342 (1998 [‘1997’]). – Codonoboea corneri (Kiew) Kiew

  • plantnameHenckelia corniculata (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 18 (1827). – Codonoboea corniculata (Jack) D.J.Middleton

  • plantnameHenckelia corrugata Mendum, Edinburgh J. Bot. 58(3): 439 (2001). – Codonoboea corrugata (Mendum) D.J.Middleton

  • plantnameHenckelia craspedodroma (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 342 (1998 [‘1997’]). – Codonoboea craspedodroma (Kiew) Kiew

  • plantnameHenckelia crenata (Baker) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Codonoboea crenata (Baker) A.Weber

  • plantnameHenckelia crinita (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 18 (1827). – Codonoboea crinita (Jack) C.L.Lim

  • plantnameHenckelia crocea (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Codonoboea crocea (Ridl.) C.L.Lim

  • plantnameHenckelia curtisii (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Codonoboea curtisii (Ridl.) C.L.Lim

  • plantnameHenckelia davisonii (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Codonoboea davisonii (Kiew) Kiew

  • plantnameHenckelia dawnii (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Codonoboea dawnii (Kiew) Kiew

  • plantnameHenckelia densifolia (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Codonoboea densifolia (Ridl.) C.L.Lim

  • plantnameHenckelia dentata (Ridl.) B.L.Burtt, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Codonoboea dentata (Ridl.) A.Weber

  • plantnameHenckelia descendens (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 343 (1998 [‘1997’]). – Codonoboea descendens (B.L.Burtt) A.Weber

  • plantnameHenckelia dibangensis (B.L.Burtt, S.K.Srivast. & Mehrotra) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia dielsii (Borza) D.J.Middleton & Mich.Möller, Taxon 60(5): 1525 (2011).

  • plantnameHenckelia diffusa B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 437 (1996). – Synonym of Loxocarpus violoides (C.B.Clarke) T.L.Yao

  • plantnameHenckelia dimidiata (Wall. ex C.B.Clarke) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia doryphylla (B.L.Burtt) A.Weber, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Codonoboea doryphylla (B.L.Burtt) C.L.Lim

  • plantnameHenckelia elegans (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Codonoboea elegans (C.B.Clarke) A.Weber

  • plantnameHenckelia elongata (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 18 (1827). – Didissandra elongata (Jack) C.B.Clarke

  • plantnameHenckelia ericiflora (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Codonoboea ericiflora (Ridl.) Ridl.

  • plantnameHenckelia ericii A.Weber, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Loxocarpus holttumii M.R.Hend.

  • plantnameHenckelia falcata (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Codonoboea falcata (Kiew) Kiew

  • plantnameHenckelia fasciata (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Codonoboea fasciata (Ridl.) C.L.Lim

  • plantnameHenckelia fasciculiflora (W.T.Wang) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia filicalyx B.L.Burtt, Thai Forest Bull., Bot. 29: 95 (2001). – Codonoboea filicalyx (B.L.Burtt) D.J.Middleton

  • plantnameHenckelia fischeri (Gamble) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]).

  • plantnameHenckelia flava (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Codonoboea flava (Ridl.) Kiew

  • plantnameHenckelia flavescens (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Codonoboea flavescens (Ridl.) Kiew

  • plantnameHenckelia flavobrunnea (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]). – Codonoboea flavobrunnea (Ridl.) Kiew

  • plantnameHenckelia floccosa (Thwaites) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 344 (1998 [‘1997’]).

  • plantnameHenckelia floribunda (M.R.Hend.) A.Weber, Beitr. Biol. Pflanzen 70: 345 (1998 [‘1997’]). – Codonoboea floribunda (M.R.Hend.) C.L.Lim

  • plantnameHenckelia follicularis (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 345 (1998 [‘1997’]). – Codonoboea follicularis (C.B.Clarke) A.Weber

  • plantnameHenckelia forrestii (J.Anthony) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia frutescens (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 18 (1827). – Didissandra frutescens (Jack) C.B.Clarke

  • plantnameHenckelia fruticola (H.W.Li) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia gambleana (C.E.C.Fisch.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 345 (1998 [‘1997’]).

  • plantnameHenckelia gardneri B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 437 (1996). – Synonym of Loxocarpus repens B.L.Burtt

  • plantnameHenckelia geitleri (A.Weber) A.Weber, Beitr. Biol. Pflanzen 70: 345 (1998 [‘1997’]). – Codonoboea geitleri (A.Weber) C.L.Lim

  • plantnameHenckelia glabrata (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 345 (1998 [‘1997’]). – Codonoboea glabrata (Ridl.) Kiew

  • plantnameHenckelia gracilipes (C.B.Clarke) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Codonoboea gracilipes (C.B.Clarke) A.Weber

  • plantnameHenckelia grandiflora (Retz.) A.Dietr., Sp. Pl., ed. 6, 1: 573 (1831). – Synonym of Lindernia antipoda (L.) Alston

  • plantnameHenckelia grandifolia A.Dietr., Sp. Pl., ed. 6, 1: 576 (1831).

  • plantnameHenckelia heterophylla (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 346 (1998 [‘1997’]). – Codonoboea heterophylla (Ridl.) C.L.Lim

  • plantnameHenckelia heterostigma (B.L.Burtt) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia hirsuta (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 346 (1998 [‘1997’]). – Codonoboea hirsuta (Ridl.) C.L.Lim

  • plantnameHenckelia hirta (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 346 (1998 [‘1997’]). – Codonoboea hirta (Ridl.) Kiew

  • plantnameHenckelia hispida (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 346 (1998 [‘1997’]). – Codonoboea hispida (Ridl.) Kiew

  • plantnameHenckelia holttumii (M.R.Hend.) A.Weber, Beitr. Biol. Pflanzen 70: 346 (1998 [‘1997’]). – Codonoboea holttumii (M.R.Hend.) C.L.Lim

  • plantnameHenckelia hookeri (C.B.Clarke) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia humboldtiana (Gardner) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 346 (1998 [‘1997’]).

  • plantnameHenckelia humilis (Miq.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 347 (1998 [‘1997’]). – Codonoboea humilis (Miq.) A.Weber

  • plantnameHenckelia inaequalis (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 347 (1998 [‘1997’]). – Codonoboea inaequalis (Ridl.) Kiew

  • plantnameHenckelia incana (Vahl) Spreng., Syst. Veg., ed. 16, 1: 38 (1824).

  • plantnameHenckelia infundibuliformis (W.T.Wang) D.J.Middleton & Mich.Möller, Taxon 60(3): 775 (2011).

  • plantnameHenckelia innominata (B.L.Burtt) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 347 (1998 [‘1997’]).

  • plantnameHenckelia insignis (C.B.Clarke) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia johorica (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 347 (1998 [‘1997’]). – Codonoboea johorica (Ridl.) Kiew

  • plantnameHenckelia kelantanensis Kiew, Gard. Bull. Singapore 61(1): 73 (2009). – Codonoboea kelantanensis (Kiew) Kiew

  • plantnameHenckelia kjellbergii B.L.Burtt, Beitr. Biol. Pflanzen 70: 378 (1998 [‘1997’]). – Codonoboea kjellbergii (B.L.Burtt) Karton. [Kartonegoro, Reference Kartonegoro2012].

  • plantnameHenckelia koerperi (B.L.Burtt) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 347 (1998 [‘1997’]). – Codonoboea koerperi (B.L.Burtt) A.Weber

  • plantnameHenckelia kolokensis B.L.Burtt, Thai Forest Bull., Bot. 29: 95 (2001). – Codonoboea kolokensis (B.L.Burtt) D.J.Middleton

  • plantnameHenckelia kompsoboea (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 347 (1998 [‘1997’]). – Codonoboea kompsoboea (C.B.Clarke) A.Weber

  • plantnameHenckelia lacei (W.W.Sm.) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia lachenensis (C.B.Clarke) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia lanceolata (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Codonoboea lanceolata (C.B.Clarke) A.Weber

  • plantnameHenckelia lancifolia (M.R.Hend.) A.Weber, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Codonoboea lancifolia (M.R.Hend.) C.L.Lim

  • plantnameHenckelia leiophylla (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Codonoboea leiophylla (Kiew) C.L.Lim

  • plantnameHenckelia leptocalyx (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Codonoboea leptocalyx (C.B.Clarke) A.Weber

  • plantnameHenckelia leucantha (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Codonoboea leucantha (Kiew) Kiew

  • plantnameHenckelia leucocodon (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Codonoboea leucocodon (Ridl.) Ridl.

  • plantnameHenckelia lilacina (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 348 (1998 [‘1997’]). – Codonoboea lilacina (Ridl.) Ridl.

  • plantnameHenckelia longipedicellata (B.L.Burtt) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia longipes (C.B.Clarke) A.Weber, Beitr. Biol. Pflanzen 70: 349 (1998 [‘1997’]). – Codonoboea longipes (C.B.Clarke) Kiew

  • plantnameHenckelia longipetiolata (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 349 (1998 [‘1997’]). – Synonym of Loxocarpus stapfii (Kraenzl.) B.L.Burtt

  • plantnameHenckelia longisepala (H.W.Li) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia lyrata (Wight) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 349 (1998 [‘1997’]).

  • plantnameHenckelia macrophylla (D.Don) Spreng., Syst. Veg., ed. 16, 4(2): 14 (1827). – Didymocarpus macrophyllus D.Don

  • plantnameHenckelia macrostachya (E.Barnes) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 349 (1998 [‘1997’]).

  • plantnameHenckelia malayana (Hook.f.) A.Weber, Beitr. Biol. Pflanzen 70: 349 (1998 [‘1997’]). – Codonoboea malayana (Hook.f.) Kiew

  • plantnameHenckelia marginata (C.B.Clarke) A.Weber, Beitr. Biol. Pflanzen 70: 349 (1998 [‘1997’]). – Codonoboea marginata (C.B.Clarke) C.L.Lim

  • plantnameHenckelia meeboldii (W.W.Sm. & Ramaswami) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 349 (1998 [‘1997’]).

  • plantnameHenckelia meijeri (B.L.Burtt) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Loxocarpus meijeri B.L.Burtt

  • plantnameHenckelia miniata (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Codonoboea miniata (Kiew) C.L.Lim

  • plantnameHenckelia minima (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Synonym of Loxocarpus incanus R.Br.

  • plantnameHenckelia mishmiensis (Debb. ex Biswas) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia missionis (Wall. ex R.Br.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]).

  • plantnameHenckelia modesta (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Codonoboea modesta (Ridl.) Kiew

  • plantnameHenckelia monantha (W.T.Wang) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia monophylla (C.B.Clarke) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia moonii (Gardner) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia murutorum (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Codonoboea murutorum (B.L.Burtt) A.Weber

  • plantnameHenckelia myricifolia (Ridl.) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Codonoboea myricifolia (Ridl.) A.Weber

  • plantnameHenckelia nana A.Weber, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Codonoboea pumila (Ridl.) C.L.Lim

  • plantnameHenckelia nervosa (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 350 (1998 [‘1997’]). – Codonoboea nervosa (C.B.Clarke) A.Weber

  • plantnameHenckelia nitida (Kiew & A.Weber) A.Weber, Beitr. Biol. Pflanzen 70: 351 (1998 [‘1997’]). – Codonoboea nitida (Kiew & A.Weber) Kiew

  • plantnameHenckelia nivea (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 351 (1998 [‘1997’]). – Codonoboea nivea Kiew

  • plantnameHenckelia oblonga (D.Don) Spreng., Syst. Veg., ed. 16, 4(2): 13 (1827). – Didymocarpus oblongus D.Don

  • plantnameHenckelia oblongifolia (Roxb.) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia oppositifolia (Retz.) A.Dietr., Sp. Pl., ed. 6, 1: 572 (1831). – Synonym of Lindernia hyssopioides (L.) Haines

  • plantnameHenckelia ovalifolia (Wight) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 351 (1998 [‘1997’]).

  • plantnameHenckelia pagonensis B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 437 (1996). – Codonoboea pagonensis (B.L.Burtt) A.Weber

  • plantnameHenckelia papillosa (M.R.Hend.) A.Weber, Beitr. Biol. Pflanzen 70: 351 (1998 [‘1997’]). Notes: Banka & Kiew (Reference Banka and Kiew2009) and Yao (Reference Yao2012) excluded Henckelia papillosa from Loxocarpus and said its affinities are with the species now placed in Codonoboea. However, Kiew & Lim (Reference Kiew and Lim2011) did not refer to the species in their Codonoboea paper. The status of the name, therefore, remains uncertain.

  • plantnameHenckelia parviflora (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 352 (1998 [‘1997’]). – Codonoboea parviflora (Ridl.) Kiew

  • plantnameHenckelia pauziana Kiew, Gard. Bull. Singapore 61(1): 74 (2009). – Codonoboea pauziana (Kiew) Kiew

  • plantnameHenckelia pectinata (Oliv.) A.Weber, Beitr. Biol. Pflanzen 70: 352 (1998 [‘1997’]). – Codonoboea pectinata (Oliv.) Kiew

  • plantnameHenckelia peduncularis (B.L.Burtt) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia petiolaris (C.B.Clarke) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Synonym of Loxocarpus verbeniflos (C.B.Clarke) B.L.Burtt

  • plantnameHenckelia platypus (C.B.Clarke) A.Weber, Beitr. Biol. Pflanzen 70: 352 (1998 [‘1997’]). – Codonoboea platypus (C.B.Clarke) C.L.Lim

  • plantnameHenckelia pleuropogon (B.L.Burtt) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Codonoboea pleuropogon (B.L.Burtt) A.Weber

  • plantnameHenckelia plicata (D.Don) Spreng., Syst. Veg., ed. 16, 4(2): 14 (1827). – Synonym of Didymocarpus macrophyllus D.Don

  • plantnameHenckelia polyanthoides (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 352 (1998 [‘1997’]). – Codonoboea polyanthoides (Kiew) C.L.Lim

  • plantnameHenckelia porphyrea B.L.Burtt, Thai Forest Bull., Bot. 29: 96 (2001). – Codonoboea porphyrea (B.L.Burtt) D.J.Middleton

  • plantnameHenckelia primulacea (C.B.Clarke) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia primulifolia (D.Don) Spreng., Syst. Veg., ed. 16, 4(2): 14 (1827). – Didymocarpus primulifolius D.Don

  • plantnameHenckelia primulina (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 352 (1998 [‘1997’]). – Codonoboea primulina (Ridl.) Kiew

  • plantnameHenckelia procumbens B.L.Burtt, Beitr. Biol. Pflanzen 70: 353 (1998 [‘1997’]). – Loxocarpus repens B.L.Burtt

  • plantnameHenckelia puerensis (Y.Y.Qian) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia pulchella (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 353 (1998 [‘1997’]). – Codonoboea pulchella (Ridl.) C.L.Lim

  • plantnameHenckelia pumila (D.Don) A.Dietr., Sp. Pl., ed. 6, 1: 574 (1831).

  • plantnameHenckelia punctata (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 353 (1998 [‘1997’]). – Codonoboea punctata (C.B.Clarke) A.Weber

  • plantnameHenckelia puncticulata (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 353 (1998 [‘1997’]). – Codonoboea puncticulata (Ridl.) C.L.Lim

  • plantnameHenckelia pycnantha (W.T.Wang) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia pyroliflora (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 353 (1998 [‘1997’]). – Codonoboea pyroliflora (Ridl.) Kiew

  • plantnameHenckelia quinquevulnera (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 353 (1998 [‘1997’]). – Codonoboea quinquevulnera (Ridl.) C.L.Lim

  • plantnameHenckelia racemosa (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 13 (1827). – Codonoboea racemosa (Jack) A.Weber

  • plantnameHenckelia ramosa (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 353 (1998 [‘1997’]). – Codonoboea ramosa (Ridl.) Kiew

  • plantnameHenckelia repens (Bedd.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 354 (1998 [‘1997’]).

  • plantnameHenckelia reptans (Jack) Spreng., Syst. Veg., ed. 16, 4(2): 14 (1827). – Codonoboea reptans (Jack) C.L.Lim

  • plantnameHenckelia reptans var. monticola (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 354 (1998 [‘1997’]). – Codonoboea reptans var. monticola (Ridl.) C.L.Lim

  • plantnameHenckelia reticulosa (C.B.Clarke) A.Weber, Beitr. Biol. Pflanzen 70: 354 (1998 [‘1997’]). – Codonoboea reticulosa (C.B.Clarke) A.Weber

  • plantnameHenckelia rexii (Hook.) Heynh., Nomencl. Bot. Hort. 1: 390 (1840). – Streptocarpus rexii (Hook.) Lindl.

  • plantnameHenckelia ridleyana A.Weber, Beitr. Biol. Pflanzen 70: 354 (1998 [‘1997’]). – Codonoboea ridleyana (B.L.Burtt) Kiew

  • plantnameHenckelia robinsonii (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 354 (1998 [‘1997’]). – Codonoboea robinsonii (Ridl.) Kiew

  • plantnameHenckelia rotundata (Barnett) D.J.Middleton & Mich.Möller, comb. nov. – Chirita rotundata Barnett, Nat. Hist. Bull. Siam Soc. 20: 16 (1961). – Type: Thailand, Chiang Mai, Me Chem, 11 vii 1922, Kerr 6274 (lecto K, designated by Barnett (Reference Barnett1961b); iso ABD, BKF, BM). This combination was included in Weber et al. (Reference Weber, Middleton, Forrest, Kiew, Lim and Rafidah2011a) but the basionym was cited as having been published in Barnett (Reference Barnett1961b) rather than the slightly earlier Barnett (Reference Barnett1961a).

  • plantnameHenckelia roxburghiana A.Dietr., Sp. Pl., ed. 6, 1: 572 (1831). – Synonym of Lindernia ruellioides (Colsm.) Pennell

  • plantnameHenckelia rubiginosa (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 355 (1998 [‘1997’]). – Codonoboea rubiginosa (Ridl.) C.L.Lim

  • plantnameHenckelia rufescens (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 355 (1998 [‘1997’]). – Loxocarpus rufescens (C.B.Clarke) B.L.Burtt

  • plantnameHenckelia rugosa (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 355 (1998 [‘1997’]). – Codonoboea rugosa (Ridl.) C.L.Lim

  • plantnameHenckelia salicina (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 355 (1998 [‘1997’]). – Codonoboea salicina (Ridl.) C.L.Lim

  • plantnameHenckelia salicinoides (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 355 (1998 [‘1997’]). – Codonoboea salicinoides (Kiew) C.L.Lim

  • plantnameHenckelia scabrinervia (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 355 (1998 [‘1997’]). – Codonoboea scabrinervia (C.B.Clarke) A.Weber

  • plantnameHenckelia scortechinii (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – The current status of this name is unknown (see Kiew & Lim (Reference Kiew and Lim2011) for further discussion)

  • plantnameHenckelia sekayuensis Banka & Kiew, Edinburgh J. Bot. 66(2): 254 (2009). – Loxocarpus incanus var. sekayuensis (Banka & Kiew) T.L.Yao

  • plantnameHenckelia semitorta (C.B.Clarke) A.Weber, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Loxocarpus semitortus (C.B.Clarke) Ridl.

  • plantnameHenckelia sericea (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Loxocarpus sericeus (Ridl.) B.L.Burtt

  • plantnameHenckelia sericiflava Kiew & Banka, Edinburgh J. Bot. 66(2): 256 (2009). – Loxocarpus sericiflavus (Kiew & Banka) T.L.Yao

  • plantnameHenckelia serrata (R.Br.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Codonoboea serrata (R.Br.) A.Weber

  • plantnameHenckelia serratifolia (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Codonoboea serratifolia (Ridl.) Kiew

  • plantnameHenckelia shuii (Z.Yu Li) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia simplex (Kraenzl.) B.L.Burtt, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Codonoboea simplex (Kraenzl.) A.Weber

  • plantnameHenckelia smitinandii (B.L.Burtt) D.J.Middleton & Mich.Möller, Taxon 60(3): 776 (2011).

  • plantnameHenckelia soldanella (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Codonoboea soldanella (Ridl.) C.L.Lim

  • plantnameHenckelia speciosa (Kurz) D.J.Middleton & Mich.Möller, Taxon 60(3): 777 (2011).

  • plantnameHenckelia stapfii (Kraenzl.) B.L.Burtt, Beitr. Biol. Pflanzen 70: 356 (1998 [‘1997’]). – Loxocarpus stapfii (Kraenzl.) B.L.Burtt

  • plantnameHenckelia stenophylla A.Weber, Beitr. Biol. Pflanzen 70: 357 (1998 [‘1997’]). – Loxocarpus angustifolius Ridl.

  • plantnameHenckelia stolonifera (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 357 (1998 [‘1997’]). – Codonoboea stolonifera (Kiew) Kiew

  • plantnameHenckelia taeniophylla B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Loxocarpus taeniophyllus (B.L.Burtt) T.L.Yao

  • plantnameHenckelia tahanica (B.L.Burtt) A.Weber, Beitr. Biol. Pflanzen 70: 357 (1998 [‘1997’]). – Codonoboea grandifolia (Ridl.) Kiew

  • plantnameHenckelia teres (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 357 (1998 [‘1997’]). – Codonoboea teres (C.B.Clarke) A.Weber

  • plantnameHenckelia tibetica (Franch.) D.J.Middleton & Mich.Möller, Taxon 60(3): 777 (2011).

  • plantnameHenckelia tiumanica (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 357 (1998 [‘1997’]). – Codonoboea tiumanica (Ridl.) C.L.Lim

  • plantnameHenckelia tunkui (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 357 (1998 [‘1997’]). – Loxocarpus tunkui Kiew

  • plantnameHenckelia urticifolia (D.Don) A.Dietr., Sp. Pl., ed. 6, 1: 574 (1831).

  • plantnameHenckelia urticoides A.Weber, Beitr. Biol. Pflanzen 70: 358 (1998 [‘1997’]). – Codonoboea urticoides (A.Weber) Kiew

  • plantnameHenckelia venusta (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 358 (1998 [‘1997’]). – Codonoboea venusta (Ridl.) Kiew

  • plantnameHenckelia verbeniflos (C.B.Clarke) B.L.Burtt, Beitr. Biol. Pflanzen 70: 358 (1998 [‘1997’]). – Loxocarpus verbeniflos (C.B.Clarke) B.L.Burtt

  • plantnameHenckelia villosa (D.Don) Spreng., Syst. Veg., ed. 16, 4(2): 13 (1827). – Didymocarpus villosus D.Don

  • plantnameHenckelia violoides (C.B.Clarke) B.L.Burtt in Coode et al., Checkl. Fl. Pl. Gymnosperms Brunei Darussalam 438 (1996). – Loxocarpus violoides (C.B.Clarke) T.L.Yao

  • plantnameHenckelia virginea (B.L.Burtt) B.L.Burtt, Beitr. Biol. Pflanzen 70: 358 (1998 [‘1997’]). – Codonoboea virginea (B.L.Burtt) A.Weber

  • plantnameHenckelia viscida (Ridl.) A.Weber, Beitr. Biol. Pflanzen 70: 359 (1998 [‘1997’]). – Codonoboea viscida (Ridl.) Kiew

  • plantnameHenckelia vulcanica (Ridl.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 359 (1998 [‘1997’]). – Codonoboea vulcanica (Ridl.) A.Weber

  • plantnameHenckelia walkerae (Gardner) D.J.Middleton & Mich.Möller, Taxon 60(3): 777 (2011).

  • plantnameHenckelia wallichiana A.Dietr., Sp. Pl., ed. 6, 1: 574 (1831). – Synonym of Henckelia urticifolia (D.Don.) A.Dietr.

  • plantnameHenckelia wightii (C.B.Clarke) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 359 (1998 [‘1997’]).

  • plantnameHenckelia woodii (Merr.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 359 (1998 [‘1997’]). – Codonoboea woodii (Merr.) A.Weber

  • plantnameHenckelia yongii (Kiew) A.Weber, Beitr. Biol. Pflanzen 70: 359 (1998 [‘1997’]). – Codonoboea yongii (Kiew) C.L.Lim

  • plantnameHenckelia zeylanica (R.Br.) A.Weber & B.L.Burtt, Beitr. Biol. Pflanzen 70: 359 (1998 [‘1997’]).

Appendix 2

A new combination in Oreocharis

para1Many new combinations in Oreocharis were published by Möller et al. (Reference Möller, Middleton, Nishii, Wei, Sontag and Weber2011b). It has come to our attention that the combination Oreocharis muscicola in that paper was not validly made. We validate it here.

  • plantnameOreocharis muscicola (Diels) Mich.Möller & A.Weber, comb. nov. – Didissandra muscicola Diels, Notes Roy. Bot. Gard. Edinburgh 5: 225 (1912). – Briggsia muscicola (Diels) Craib, Notes Roy. Bot. Gard. Edinburgh 11: 264 (1919). – Type: China, Yunnan, Mekong-Salwin [Salween] divide, vii 1905, Forrest 5095 (holo E). – Distribution: SW China, Bhutan, NE India, Burma.

References

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