Connectivity at the land–water interface

doi:10.1017/CBO9780511754821.006

5 - Connectivity at the land–water interface

Published online by Cambridge University Press: 24 May 2010

Drew M. Talley ,

Gary R. Huxel and

Marcel Holyoak

Edited by

Kevin R. Crooks and

M. Sanjayan

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Kevin R. Crooks: Affiliation:
Colorado State University
M. Sanjayan: Affiliation:
The Nature Conservancy, Virginia

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Summary

INTRODUCTION

There is a growing appreciation in ecology and conservation that even those habitats and ecosystems (we will use these terms interchangeably) traditionally considered “insular” are in fact reticulately connected (Polis and Strong 1996). These connections are mediated by both physical and biological processes spanning a wide range of spatial and temporal scales (Polis and Strong 1996). This is clearly true of linkages between similar habitat types, such as fragmented systems, or patchily distributed communities (e.g., see Moilanen and Hanski Chapter 3; DiBacco et al. Chapter 8). However, linkages are also critically important between very distinct habitats, such as forests and grasslands, freshwater and marine habitats, or aquatic and terrestrial ecosystems. In fact, as we will discuss in this chapter, if connected habitats are dissimilar we believe there is at least as much potential for movement of materials to influence the connected systems as when habitats are similar.

Inter-habitat connectivity has been formally recognized at least since 1923 (Summerhayes and Elton 1923), but recognition of the importance of linking systems for altering their productivity goes back much further (e.g., to ancient irrigation schemes, such as those in the Nile Delta of Egypt). Recently, an integration of the landscape and ecosystem ecology concepts of connectivity, and their potential to structure ecosystems, has begun to be examined in detail (e.g., Polis and Hurd 1996).

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Type: Chapter
Information: Connectivity Conservation , pp. 97 - 129

DOI: https://doi.org/10.1017/CBO9780511754821.006 [Opens in a new window]

Publisher: Cambridge University Press

Print publication year: 2006

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References

Anderson, A. M., Haukos, D. A., and Anderson, J. T.. 1999. Diet composition of three anurans from the playa wetlands of northwest Texas. Copeia 1999:515–520CrossRef Google Scholar

Anderson, W. B., and Polis, G. A.. 1999. Nutrient fluxes from water to land: seabirds affect plant nutrient status on Gulf of California islands. Oecologia 118:324–332CrossRef Google Scholar PubMed

Anderson, W. B., and Wait, D. A.. 2001. Subsidized island biogeography hypothesis: another new twist on an old theory. Ecology Letters 4:289–291CrossRef Google Scholar

Arndt, R. G. 1999. Predation by the black iguana (Ctenosaura similis) on the painted ghost crab (Ocypode gaudichaudii) in Costa Rica. Florida Scientist 62:111–114Google Scholar

Barrett, K., Wait, D. A., and Anderson, W. B.. 2003. Small island biogeography in the Gulf of California: lizards, the subsidized island biogeography hypothesis, and the small island effect. Journal of Biogeography 30:1575–1581CrossRef Google Scholar

Blanchard, D. C., and Syzdek, L.. 1970. Mechanism for the water-to-air transfer and concentration of bacteria. Science 170:626–628CrossRef Google Scholar

Bodkin, J. L., and Jameson, R. J.. 1991. Patterns of seabird and mammal carcass deposition along the central California coast, 1980–1986. Canadian Journal of Zoology 69:1149–1155CrossRef Google Scholar

Bosman, A. L., and Hockey, P. A. R.. 1988. The influence of seabird guano on the biological structure of rocky intertidal communities on islands off the west coast of southern Africa. South African Journal of Marine Science 7:61–68CrossRef Google Scholar

Cadenasso, M. L., Pickett, S. T. A., Weathers, K. C., and Jones, C. G.. 2003. A framework for a theory of ecological boundaries. BioScience 53:750–758CrossRef Google Scholar

Carlton, J. T., and Hodder, J.. 2003. Maritime mammals: terrestrial mammals as consumers in marine intertidal communities. Marine Ecology–Progress Series 256:271–286CrossRef Google Scholar

Carpenter, S. R., Ludwig, D., and Brock, W. A.. 1999. Management of eutrophication for lakes subject to potentially irreversible change. Ecological Applications 9:751–771CrossRef Google Scholar

Carr, M. H., Neigel, J. E., Estes, J. A., et al. 2003. Comparing marine and terrestrial ecosystems: implications for the design of coastal marine reserves. Ecological Applications 13:S90–S107CrossRef Google Scholar

Cheng, L. (ed.) 1976. Marine Insects. Amsterdam, Netherlands: North Holland.Google Scholar

Collier, K. J., Bury, S., and Gibbs, M.. 2002. A stable isotope study of linkages between stream and terrestrial food webs through spider predation. Freshwater Biology 47:1651–1659CrossRef Google Scholar

Cortes, J., and Risk, M. J.. 1985. A reef under siltation stress: Cahuita, Costa Rica. Bulletin of Marine Science 36:339–356Google Scholar

Crowder, L. B., Squires, D. D., and Rice, J. A.. 1997. Nonadditive effects of terrestrial and aquatic predators on juvenile estuarine fish. Ecology 78:1796–1804CrossRef Google Scholar

Cummins, K. W., Petersen, R. C., Howard, F. O., Wuycheck, J. C., and Holt, V. I.. 1973. The utilization of leaf litter by stream detritivores. Ecology 54:336–345CrossRef Google Scholar

Cyrus, D. P., Martin, T. J., and Reavell, P. E.. 1997. Salt-water intrusion from the Mzingazi River and its effects on adjacent swamp forest at Richards Bay, Zululand, South Africa. Water SA 23:101–108Google Scholar

Dahm, C. N., Cummins, K. W., Valett, H. M., and Coleman, R. L.. 1995. An ecosystem view of the restoration of the Kissimmee River. Restoration Ecology 3:225–238CrossRef Google Scholar

Daniels, R. B. 1960. Entrenchment of the Willow drainage ditch. American Journal of Science 258:167–176CrossRef Google Scholar

Darimont, C. T., Reimchen, T. E., and Paquet, P. C.. 2003. Foraging behaviour by gray wolves on salmon streams in coastal British Columbia. Canadian Journal of Zoology–Revue Canadienne de Zoologie 81:349–353CrossRef Google Scholar

Dayton, P. K., Thrush, S. F., Agardy, M. T., and Hofman, R. J.. 1995. Environmental effects of marine fishing. Aquatic Conservation: Marine and Freshwater Ecosystems 5:205–232CrossRef Google Scholar

Dayton, P. K., Thrush, S., and Coleman, F. C.. 2002. Ecological Effects of Fishing in Marine Ecosystems of the United States. Arlington, VA: Pew Oceans Commission.Google Scholar

DeAngelis, D. L. 1992. Dynamics of Nutrient Cycling and Food Webs. London: Chapman and Hall.CrossRef Google Scholar

DeMaster, D. P., Fowler, C. W., Perry, S. L., and Richlen, M. E.. 2001. Predation and competition: the impact of fisheries on marine-mammal populations over the next one hundred years. Journal of Mammalogy 82:641–6512.0.CO;2>CrossRef Google Scholar

Dias, P. C. 1996. Sources and sinks in population biology. Trends in Ecology and Evolution 11:326–330CrossRef Google Scholar PubMed

DiBacco, C., and Chadwick, D. B.. 2001. Assessing the dispersal and exchange of brachyuran larvae between regions of San Diego Bay, California and nearshore coastal habitats using elemental fingerprinting. Journal of Marine Research 59:53–78CrossRef Google Scholar

Farina, J. M., Salazar, S., Wallem, K. P., Witman, J. D., and Ellis, J. C.. 2003. Nutrient exchanges between marine and terrestrial ecosystems: the case of the Galapagos sea lion Zalophus wollebaecki. Journal of Animal Ecology 72:873–887CrossRef Google Scholar

Forman, R. T. T., and Alexander, L. E.. 1998. Roads and their major ecological effects. Annual Review of Ecology and Systematics 29:207–231CrossRef Google Scholar

Gagliano, S. M., Meyerarendt, K. J., and Wicker, K. M.. 1981. Land loss in Mississippi River deltaic plain. Transaction of the Gulf Coast Association of the Geological Societies 31:295–300Google Scholar

Gasith, A., and Hasler, A. D.. 1976. Airborne litterfall as a source of organic matter in lakes. Limnology and Oceanography 21:253–258CrossRef Google Scholar

Gende, S. M., Edwards, E. D., Willson, M. F., and Wipfli, M. S.. 2002. Pacific salmon in aquatic and terrestrial ecosystems. BioScience 52:917–928CrossRef Google Scholar

Gerlanc, N. M., and Kaufman, G. A.. 2003. Use of bison wallows by anurans on Konza Prairie. American Midland Naturalist 150:158–168CrossRef Google Scholar

Gillanders, B. M., Able, K. W., Brown, J. A., Eggleston, D. B., and Sheridan, P. F.. 2003. Evidence of connectivity between juvenile and adult habitats for mobile marine fauna: an important component of nurseries. Marine Ecology–Progress Series 247:281–295CrossRef Google Scholar

Gray, L. J. 1993. Response of insectivorous birds to emerging aquatic insects in riparian habitats of a tallgrass prairie stream. American Midland Naturalist 129:288–300CrossRef Google Scholar

Griffin, D. W., Kellogg, C. A., Garrison, V. H., and Shinn, E. A.. 2002. The global transport of dust: an intercontinental river of dust, microorganisms and toxic chemicals flows through the Earth's atmosphere. American Scientist 90:228–235CrossRef Google Scholar

Grimm, N. B. 1987. Nitrogen dynamics during succession in a desert stream. Ecology 68:1157–1170CrossRef Google Scholar

Hansen, A. J., and di Castri, F. (eds.) 1992. Landscape Boundaries: Consequences for Biotic Diversity, and Ecological Flows. New York: Springer-Verlag.CrossRef Google Scholar

Hanski, I. A., and Gilpin, M. E. (eds.) 1997. Metapopulation Biology: Ecology, Genetics, and Evolution. San Diego, CA: Academic Press.Google Scholar

Harvell, C. D., Kim, K., Burkholder, J. M., et al. 1999. Review: Marine ecology–emerging marine diseases–climate links and anthropogenic factors. Science 285:1505–1510CrossRef Google Scholar

Hemminga, M. A., and Nieuwenhuize, J.. 1990. Seagrass wrack-induced dune formation on a tropical coast (Banc d'Arguin, Mauritania). Estuarine, Coastal and Shelf Science 31:499–502CrossRef Google Scholar

Hewitt, J. E., Cummings, V. J., Ellis, J. I., et al. 2003. The role of waves in the colonisation of terrestrial sediments deposited in the marine environment. Journal of Experimental Marine Biology and Ecology 290:19–47CrossRef Google Scholar

Huxel, G. R., and McCann, K.. 1998. Food web stability: the influence of trophic flows across habitats. American Naturalist 152:460–469CrossRef Google Scholar PubMed

Huxel, G. R., McCann, K., and Polis, G. A.. 2002. Effects of partitioning allochthonous and autochthonous resources on food web stability. Ecological Research 17:419–432CrossRef Google Scholar

Huxel, G. R., G. A. Polis, and R. D. Holt., 2004. At the frontier of the integration of food web ecology and landscape ecology. Pp. 434–451 in Polis, G. A., Power, M. E., and Huxel, G. R. (eds.) Food Webs at the Landscape Level. Chicago, IL: University of Chicago Press.Google Scholar

Inman, D. L. 2005. Littoral cells. Pp. 594–598 in M. Schwartz (ed.) Encyclopedia of Coastal Science. New York: Springer-Verlag.

Jackson, J. K., and Fisher, S. G.. 1986. Secondary production, emergence, and export of aquatic insects of a Sonoran desert stream. Ecology 67:629–638CrossRef Google Scholar

Karr, J. R., and Schlosser, I. J.. 1978. Water resources and the land–water interface. Science 201:229–234CrossRef Google Scholar

Koebel, J. W. 1995. An historical perspective on the Kissimmee River restoration project. Restoration Ecology 3:149–159CrossRef Google Scholar

Kupferberg, S. J. 1997. Bullfrog (Rana catesbeiana) invasion of a California river: the role of larval competition. Ecology 78:1736–1751CrossRef Google Scholar

Laurance, W. H., Didham, R. K., and Power, M. E.. 2001. Ecological boundaries: a search for synthesis. Trends in Ecology and Evolution 16:70–71CrossRef Google Scholar

Levins, R. 1969. Some demographic and genetic consequences of environmental heterogeneity for biological control. Bulletin of the Entomological Society of America 15:237–240CrossRef

Likens, G. E., and Bormann, F. H.. 1974. Linkages between terrestrial and aquatic ecosystems. BioScience 24:447–456CrossRef Google Scholar

Loreau, M., Mouquet, N., and Holt, R. D.. 2003. Meta-ecosystems: a theoretical framework for a spatial ecosystem ecology. Ecology Letters 6:673–679CrossRef Google Scholar

Luo, H. R., Smith, L. M., Haukos, D. A., and Allen, B. L.. 1999. Sources of recently deposited sediments in playa wetlands. Wetlands 19:176–181CrossRef Google Scholar

Malanson, G. P. 1993. Riparian Landscapes. Cambridge, U.K: Cambridge University Press.CrossRef Google Scholar

Mann, K. H., and Lazier, R. N.. 1991. Dynamics of Marine Ecosystems. Oxford, UK: Blackwell.Google Scholar

Marra, P. P., and Holmes, R. T.. 1997. Avian removal experiments: do they test for habitat saturation or female availability?Ecology 78:947–952CrossRef Google Scholar

McCann, K., Hastings, A., and Huxel, G. R.. 1998. Weak trophic interactions and the balance of nature. Nature 395:794–798CrossRef Google Scholar

McHugh, D. J. 2003. A Guide to the Seaweed Industry. FAO Fisheries Technical Paper No. 441. Rome, Italy: Food and Agriculture Organization of the United Nations.Google Scholar

McPeek, M. A., and Kalisz, S.. 1998. On the joint evolution of dispersal and dormancy in metapopulations. Advances in Limnology 52:33–51Google Scholar

Minshall, G. W. 1967. Role of allochthonous detritus in the trophic structure of a woodland springbrook community. Ecology 48:139–149CrossRef Google Scholar

Mitsch, W. J., and Gosselink, J. G.. 2000. Wetlands, 3rd edn. New York: John Wiley.Google Scholar

Naiman, R. J., and Decamps, H.. 1997. The ecology of interfaces: riparian zones. Annual Review of Ecology and Systematics 28:621–658CrossRef Google Scholar

Naiman, R. J., Melillo, J. M., Lock, M. A., Ford, T. E., and Reice, S. R.. 1987. Longitudinal patterns of ecosystem processes and community structure in a sub-arctic river continuum. Ecology 68:1139–1156CrossRef Google Scholar

Nakano, S., and Murakami, M.. 2001. Reciprocal subsidies: dynamic interdependence between terrestrial and aquatic food webs. Proceedings of the National Academy of Sciences of the USA 98:166–170CrossRef Google Scholar PubMed

Nakano, S., Miyasaka, H., and Kuhara, N.. 1999. Terrestrial–aquatic linkages: riparian arthropod inputs alter trophic cascades in a stream food web. Ecology 80:2435–2441Google Scholar

Neal, C., Leeks, G. J. L., Millward, G. E., et al. 2003. Land–ocean interaction: processes, functioning and environmental management from a UK perspective: an introduction. Science of the Total Environment 314:3–11CrossRef Google Scholar PubMed

Nehlsen, W., Williams, J. E., and Lichatowich, J. A.. 1991. Pacific salmon at the crossroads: stocks at risk from California, Oregon, Idaho, and Washington. Fisheries 16:4–212.0.CO;2>CrossRef Google Scholar

Norkko, A., Thrush, S. F., Hewitt, J. E., et al. 2002. Smothering of estuarine sandflats by terrigenous clay: the role of wind–wave disturbance and bioturbation in site-dependent macrofaunal recovery. Marine Ecology – Progress Series 234:23–41CrossRef Google Scholar

Odum, W. E. 1980. The status of three ecosystem-level hypotheses regarding salt marsh estuaries: tidal subsidy, outwelling, and detritus-based food chains. Pp. 485–495 in Kennedy, V. S. (ed.) Estuarine Perspectives. New York: Academic Press.Google Scholar

Ohio Lake Erie Commission, . 2000. Lake Erie Protection and Restoration Plan. Toledo, OH: Ohio Lake Erie Commission. Available online at http://www.epa.state.oh.US/oleo/index.html/Google Scholar

Polis, G. A., and Hurd, S. D.. 1995. Extraordinarily high spider densities on islands: Flow of energy from the marine to terrestrial food webs and the absence of predation. Proceedings of the National Academy of Sciences of the USA 92:4382–4386CrossRef Google Scholar PubMed

Polis, G. A., and Hurd, S. D.. 1996. Linking marine and terrestrial food webs: allochthonous input from the ocean supports high secondary productivity on small islands and coastal land communities. American Naturalist 147:396–423CrossRef Google Scholar

Polis, G. A., and Strong, D. R.. 1996. Food web complexity and community dynamics. American Naturalist 147:813–846CrossRef Google Scholar

Polis, G. A., Anderson, W. B., and Holt, R. D.. 1997a. Toward an integration of landscape and food web ecology: the dynamics of spatially subsidized food webs. Annual Review of Ecology and Systematics 29:289–316CrossRef Google Scholar

Polis, G. A., Hurd, S. D., Jackson, C. T., and Piñero, F. S.. 1997b. El Niño effects on the dynamics and control of an island ecosystem in the Gulf of California. Ecology 78:1884–1897Google Scholar

Polis, G. A., Power, M. E., and Huxel, G. R. (eds.) 2004. Food Webs at the Landscape Level. Chicago, IL: University of Chicago Press.Google Scholar

Power, M. E. 1984. Depth distributions of armored catfish: predator-induced resource avoidance?Ecology 65:523–528CrossRef Google Scholar

Power, M. E. 2001. Prey exchange between a stream and its forested watershed elevates predator densities in both habitats. Proceeding of the National Academy of Sciences of the USA 98:14–15CrossRef Google Scholar PubMed

Rodriguez-Lanetty, M., and Hoegh-Guldberg, O.. 2002. The phylogeography and connectivity of the latitudinally widespread scleractinian coral Plesiastrea versipora in the Western Pacific. Molecular Ecology 11:1177–1189CrossRef Google Scholar PubMed

Roff, D. A. 2002. Life History Evolution.Sunderland, MA: Sinauer Associates.Google Scholar

Rose, M. D., and Polis, G. A.. 1998. The distribution and abundance of coyotes: the effects of allochthonous food subsidies from the sea. Ecology 79:998–1007CrossRef Google Scholar

Rosenzweig, M. L. 1991. Habitat selection and population interactions: - the search for mechanism. American Naturalist 137:S5–S28CrossRef Google Scholar

Runyan, K., and Griggs, G. B.. 2003. The effects of armoring seacliffs on the natural sand supply to the beaches of California. Journal of Coastal Research 19:336–347Google Scholar

Sabo, J. L., and Power, M. E.. 2002. Numerical response of lizards to aquatic insects and short-term consequences for terrestrial prey. Ecology 83:3023–3036CrossRef Google Scholar

Sánchez-Piñero, F., and Polis, G. A.. 2000. Bottom–up dynamics of allochthonous input: direct and indirect effects of seabirds on islands. Ecology 81:3117–3132Google Scholar

Sanzone, D. M., Meyer, J. L., Marti, E., et al. 2003. Carbon and nitrogen transfer from a desert stream to riparian predators. Oecologia 134:238–250CrossRef Google Scholar PubMed

Schoenherr, A. A. 1992. A Natural History of California. Berkeley, CA: University of California Press.Google Scholar

Southwood, T. R. E. 1962. Migration of terrestrial arthropods in relation to habitat. Biological Reviews 37:171–214CrossRef Google Scholar

Stapp, P., Polis, G. A., and Sanchez-Piñero, F.. 1999. Stable isotopes reveal strong marine and El Niño effects on island food webs. Nature 401:467–469CrossRef Google Scholar

Steele, J. H., S. R. Carpenter, J. E. Cohen, P. K. Dayton, and R. E. Ricklefs., 1993. Comparing terrestrial and marine ecological systems. Pp. 1–12 in Levin, S. A., Powell, T. M., and Steele, J. H. (eds.) Patch Dynamics. New York: Springer-Verlag.CrossRef Google Scholar

Strayer, D. L., Power, M. E., Fagan, W. F., Pickett, S. T. A., and Belnap, J.. 2003. A classification of ecological boundaries. BioScience 53:723–729CrossRef Google Scholar

Summerhayes, V. S., and Elton, C. S.. 1923. Contributions to the ecology of Spitzbergen and Bear Island. Journal of Ecology 11:214–286CrossRef Google Scholar

Talley, D. M., North, E. W., Juhl, A. R., et al. 2003. Research challenges at the land–sea interface. Estuarine Coastal and Shelf Science 58:699–702CrossRef Google Scholar

Talley, T. S., Crooks, J. A., and Levin, L. A.. 2001. Habitat utilization and alteration by the invasive burrowing isopod, Sphaeroma quoyanum, in California salt marshes. Marine Biology 138:561–573CrossRef Google Scholar

Thrush, S. F., Pridmore, R. D., Hewitt, J. E., and Cummings, V. J.. 1994. The importance of predators on a sandflat: interplay between seasonal changes in prey densities and predator effects. Marine Ecology – Progress Series 107:211–222CrossRef Google Scholar

Thrush, S. F., Hewitt, J. E., Cummings, V., et al. 2004. Muddy waters: elevating sediment input to coastal and estuarine habitats. Frontiers in Ecology and the Environment 2:299–306CrossRef Google Scholar

Toth, L. A. 1995. Principles and guidelines for restoration of river/floodplain ecosystems: Kissimmee River, Florida. Pp. 49–75 in J. Cairns Jr. (ed.) Rehabilitating Damaged Ecosystems. Boca Raton, FL: Lewis Publications.

Wainright, S. C., Haney, J. C., Kerr, C., Golovkin, A. N., and Flint, M. V.. 1998. Utilization of nitrogen derived from seabird guano by terrestrial and marine plants at St. Paul, Pribilof Islands, Bering Sea, Alaska. Marine Biology 131:63–71CrossRef Google Scholar

Wallace, J. B., Eggert, S. L., Meyer, J. L., and Webster, J. R.. 1997. Multiple trophic levels of a forest stream linked to terrestrial litter inputs. Science 277:102–104CrossRef Google Scholar

Ward, J. V. 1998. Riverine landscapes: biodiversity patterns, disturbance regimes, and aquatic conservation. Biological Conservation 83:269–278CrossRef Google Scholar

Watson, R., and Pauly, D.. 2001. Systematic distortions in world fisheries catch trends. Nature 414:534–536CrossRef Google Scholar PubMed

Wiens, J. A. 2002. Riverine landscapes: taking landscape ecology into the water. Freshwater Biology 47:501–515CrossRef Google Scholar

Wikelski, M., and Thom, C.. 2000. Marine iguanas shrink to survive El Niño: changes in bone metabolism enable these adult lizards to reversibly alter their length. Nature 403:37–38CrossRef Google Scholar

Wilbur, H. M. 1980. Complex life cycles. Annual Review of Ecology and Systematics 11:67–93CrossRef Google Scholar

Willson, M. F., Gende, S. M., and Marston, B. H.. 1998. Fishes and the forest: expanding perspectives on fish–wildlife interactions. BioScience 48:455–462CrossRef Google Scholar

Willson, M. F., S. M. Gende, and P. A. Bisson., 2004. Anadromous fishes as ecological links between ocean, fresh water, and land. Pp. 284–300 in Polis, G. A., Power, M. E., and Huxel, G. R. (eds.) Food Webs at the Landscape Level. Chicago, IL: University of Chicago Press.Google Scholar

Witman, J. D., J. C. Ellis, and W. B. Anderson. 2004. The influence of physical processes, organisms, and permeability on cross-ecosystem fluxes. Pp. 335–358 in Polis, G. A, Power, M. E, and Huxel, G. R (eds.) Food Webs at the Landscape Level. Chicago, IL: University of Chicago Press.Google Scholar

Wootton, J. T. 1991. Direct and indirect effects of nutrients on intertidal community structure: variable consequences of seabird guano. Journal of Experimental Marine Biology and Ecology 151:139–154CrossRef Google Scholar

Wright, J. P., Gurney, W. S. C., and Jones, C. G.. 2004. Patch dynamics in a landscape modified by ecosystem engineers. Oikos 105:336–348CrossRef Google Scholar

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